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Identification guide to the Leaf Beetles of Great Smoky Mountains National Park

Working draft, February 2004

Edward G. Riley
Department of Entomology
Texas A&M University
College Station, TX 77845


Production of this work is supported by the small grants program of Discover Life in America.

This identification guide includes many species that are not formally recorded from GSMNP but may be found there.

Species preceded by an asterisk "*" are confirmed for park.

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Go directly to Criocerinae, couplet 10
Go directly to most Eumolpinae, couplet 34
Go directly to Cryptocephalinae, couplet 12
Go directly to Chrysomelinae, couplet 29
Go directly to Cassidinae – the tortoise beetles, couplet 20
Go directly to Cassidinae – the hispine beetles, couplet 24
Go directly to Galerucinae s. str., couplet 65
Go directly to Galerucinae, Alticini, couplet 42



1a      Prothorax with raised marginal bead or sharp ridge extending for entire or nearly entire length [HELP] ... Family Chrysomelidae (large part) … 11

1b      Prothorax lacking marginal bead or ridge, or if present, weak and only evident near base [HELP] ... 2

2a      Dorsum largely hairy or scaly ... 3

2b      Dorsum not conspicuously hairy or scaly … 8

3a      Usually small compact beetle with hind femur enlarged; prothorax tapered towards anterior margin; head tucked partially below anterior margin of prothorax … Family Chrysomelidae, Subfamily Bruchinae

Acanthoscelides spp.
*Altheas hibisci
*Amblycerus robiniae
*Gibbobruchus mimus
*Stylantheus macrocerus

3b      Larger, non-compact beetle with hind femur not enlarged; prothorax subcylindrical, not especially tapered toward anterior margin; head exposed, not tucked below anterior margin of thorax … 4

4a      Prothorax strongly angled at middle; general body form elongate and parallel sided, resembling members of the family Cleridae …Family Megalopodidae, Subfamily Zeugophorinae

Zeugophora scutellaris

4b      Prothorax not or only weakly angled at middle (but may have small teeth); body form of beetle shorter and broader, not resembling members of the family Cleridae …. Family Chrysomelidae, Subfamily Eumolpinae (small part) … 5

5a      Middle and hind tibia with distal outer margin excavated and usually with a stiff brush of setae [HELP]; body dark metallic with whitish hairs

Graphops curtipennis
Graphops marcassita
Graphops pubescens

5b      Middle and hind tibia not modified as above; body usually not metallic … 6

6a      Prothorax with short row of teeth on lateral margin

*Demotina modesta
Myochrous denticollis

6b      Prothorax without teeth on lateral margin … 7

7a      Prothorax cylindrical, weakly punctate with simple punctures; anterior and posterior femur without ventral tooth

Bromius obscurus
Fidia viticida

7b      Prothorax depressed, subcylindrical, distinctly punctate with large ocellate punctures; anterior and posterior femur with minute ventral tooth

Xanthonia angulata
*Xanthonia decemnotata
Xanthonia monticola
*Xanthonia serrata
Xanthonia stevensi
*Xanthonia striata
Xanthonia villosula
*Xanthonia sp.

8a (from 2)  Head with frontal area flat and smooth, without X-shaped groove

Elytra without elevated ridges, punctation confused … Family Orsodacnidae

*Orsodacne atra

Elytra with raised ridges, punctures in rows … Family Chrysomelidae, Subfamily Eumolpinae, Tribe Synetini

*Syneta extorris
*Syneta ferruginea

8b      Head with X-shaped groove on frontal area … 9


9a      Hind femur with one or more ventral teeth …Family Chrysomelidae, Subfamily Donaciinae

Inner edge of apical portion of elytral suture exposed

Plateumaris fulvipes
*Plateumaris metallica
*Plateumaris rufa

Inner edge of apical portion of elytral suture concealed

Donacia biimpressa
*Donacia caerulea
Donacia confluenta
Donacia liebecki
Donacia subtilis

9b      Hind femur without ventral tooth …Family Chrysomelidae, Subfamily Criocerinae … 10

10a    Prothorax with distinct median constriction; claws of each tarsus fused together at base, subparallel

Elytral uniformly dark

Oulema collaris
Oulema cornuta
*Oulema maculicollis
*Oulema melanopus
Oulema palustris
*Oulema sayi

Elytral pale with dark stripes

Lema daturaphila
Lema trivittata trivittata

Elytral dark with broad pale band

Lema solani

Elytral red with black spots

*Neolema cordata
*Neolema sexpunctata

10b    Prothorax weakly constricted near base; claws of each tarsus not obviously fused at base, divergent

Crioceris asparagi
Crioceris duodecimpunctata

11a (from 1) Body form cylindrical, somewhat barrel-shaped and generally compact; face and pygidium subvertical to vertical [HELP] …12

11b    Body form generally depressed, not especially cylindrical or compact; face and pygidium not vertical … 19

12a    Compact, rough-backed beetle with legs and antennae fitting neatly into grooves and pockets on ventral surface of body; suture of elytra serrate, zipper-like … Subfamily Cryptocephalinae, Tribe Chlamisini

Smaller beetle, mostly black, non-metallic      

Chlamisus foveolatus
Exema canadensis
*Exema dispar

Larger beetle, coppery, bronze or metallic brownish

Neochlamisus assimilis
Neochlamisus bebbianae
Neochlamisus bimaculatus
Neochlamisus eubati
*Neochlamisus gibbosus
Neochlamisus platani
Neochlamisus tuberculatus

12b    Not rough-backed (but often deeply punctured), legs and antennae not fitting into grooves and pockets on ventral surface of body … 13

13a    Antenna shorter, not reaching postero-lateral angle of pronotum, and with several distal segments weakly to distinctly serrate (saw-like)… Subfamily, Cryptocephalinae, Tribe Clytrini … 18

13b    Antenna longer, nearly reaching to surpassing postero-lateral angle of pronotum, and filiform (thread-like) or with several distal segments slightly enlarged … 14

14a    Base of pronotum with distinct bead-like margin [HELP]

A. Anterior femur not enlarged, equal in size to middle femur

Pachybrachis hepaticus

AA. Anterior femur enlarged, notably larger than middle femur

B. Dorsum hairy

Pachybrachis morosus

BB. Dorsum without hairs

C. Elytron with pale or dark stripes

Pachybrachis bivittatus
Pachybrachis litigosus
Pachybrachis othonus

CC. Elytron without stripes

D. Large robust beetle, length greater than 4.7 mm, with punctation of pronotum sparse

Griburius scutellaris

DD. Smaller and less robust beetle, length less than 4.7 mm, with punctation of pronotum dense

Elytra mostly black or black with sharp, well-defined pattern of paler markings

Pachybrachis luctuosus
*Pachybrachis luridus
Pachybrachis m-nigrum
Pachybrachis nigricornis carbonarius
Pachybrachis picturatus
Pachybrachis subfasciatus
*Pachybrachis tridens
Pachybrachis trinoatus
Pachybrachis viduatus

Elytra mostly brown, yellow or reddish, usually with poorly defined markings, or black to near black with sparse pale flecks

Pachybrachis atomarius
Pachybrachis confederatus
Pachybrachis femoratus
Pachybrachis obfuscatus
*Pachybrachis obsoletus
*Pachybrachis pectoralis
Pachybrachis relictus
Pachybrachis spumarius

14b    Base of pronotum not margined [HELP]… 15


15a    Larger species, length greater than 2.5 mm … 16

15b    Smaller species, length less than 2.5 mm … 17

16a    Entirely dark metallic in color, usually dark bluish metallic, prosternum nearly as wide as long, each tarsal claw with broad basal tooth

Lexiphanes affinis
*Lexiphanes saponatus
Lexiphanes seminulum

16b    Color variable but not entirely dark bluish metallic, prosternum narrower than long, each tarsal claw without basal tooth

A. Punctation at apex of elytra deep and confused

Cryptocephalus schreibersii
Cryptocephalus tinctus

AA. Punctation at apex of elytra deep to shallow, not confused     

B. Pronotum with longitudinal stripes

Cryptocephalus leucomelas

BB. Pronotum without longitudinal stripes

Elytron entirely dark

Cryptocephalus nanus

Elytron black with red spot or spots

*Bassareus mammifer
Cryptocephalus notatus
*Cryptocephalus quadriplex
Cryptocephalus binominus

Elytron black with orange lateral stripe

Bassareus lituratus

Elytron brownish with pale spots

Bassaerus brunnipes
Bassareus clathratus
Bassaerus formosus
Cryptocephalus badius
Cryptocephalus guttulatus

Elytron pale with darker stripes

Cryptocephalus calidus
Cryptocephalus gibbicollis
Cryptocephalus incertus
*Cryptocephalus venustus

Elytron pale yellowish with irregular dark markings

Cryptocephalus mutabilis

17a (from 15) Elytron dark metallic

*Diachus auratus
Diachus pallidicornis

17b    Elytron black to pale brownish, non-metallic

Triachus atomus
*Triachus cerinus

18a (from 13) Prothorax conspicuously hairy

Coleothorpa aenescens
Coleothorpa dominicana

18b    Prothorax not hairy

Elytron dark with red humeral spot

Saxinis omogera
*Babia quadriguttata

Elytron pale yellowish or mostly pale yellowish

Anomoea flavokansiensis
*Anomoea laticlavia

19a (from 11) Margins of both elytra and pronotum expanded, most of head and legs not visible from dorsal view … Subfamily Cassidinae – the tortoise beetles … 20

19b    Margins of elytra and pronotum not both expanded, margins of pronotum may be somewhat expanded but then not covering legs and most of head … 22

20a    Head not completely covered by pronotum, narrowly visible from above; dorsum red with dark spots on elytra

Chelymorpha cassidea

20b    Head completely covered by pronotum, not visible from above; color variable … 21

21a    Large species, length greater than 8 mm; pronotum pale with one or three small black spots

*Physonota helianthi
*Physonota unipunctata

21b    Smaller species, length usually less than 8 mm; pronotum not as above

A. Dark coloration of elytral disc extended on expanded margin of elytra

Elytral surface smooth, pale

*Charidotella purpurata

Elytral surface smooth, mostly black or usually black with irregular yellow patches

*Deloyala guttata

Elytral surface rough and uneven, dark brownish

*Plagiometriona clavata clavata

AA. Expanded margins of elytra entirely pale

Disc of elytra dark with pale stripes

Agroiconota bivittata

Disc of elytra entirely pale, yellow to greenish

Gratiana pallidula

Disc of elytra pale yellowish to reddish and with small black spots

*Charidotella sexpunctata bicolor
*Jonthonota nigripes

22a (from 19) Insertion points of antennae fairly close together, high on face at a point approximately level to middle of eyes … 23

22b    Insertion points of antennae far apart, separated by most the width of the face, low on face at point approximately level with lower margins of eyes … 28

23a    Mouthparts positioned below and at the back of head, not projecting, usually partially covered by the front margin of the prosternum; antennae stiff and generally projected forward … Subfamily Cassidinae – the hispines … 24

23b    Mouthparts positioned at end of head, generally projecting, not covered partially by front margin of prosternum; antennae more flexible, projecting in various directions … Subfamily Galerucinae … 41

24a    Elytra relatively smooth and with rows of fine punctures; color black without markings

Stenispa metallica

24b    Elytra coarsely punctate, punctures in rows or not … 25

25a    Antenna composed of 11 distinct segments, not or weakly clubbed … 26


25b    Antenna apparently composed of 8 segments, the distal joints fused to form a weak to distinct club

Elytra brownish with vague darker spots

Brachycoryna pumila

Elytra dark with pale stripes

Microrhopala vittata

Microrhopala xerene

Elytra entirely dark

Glyphuroplata pluto
Microrhopala erebus
Microrhopala excavata
Microrhopala hecate
Octotoma plicatula

26a    Middle of each elytron with 10 rows of punctures

Elytra black with humeral areas orange

*Odontota mundula
Odontota scapularis

Elytra red or orange with sutural area black

*Odontota dorsalis
Odontota horni

Elytra orange with darker brownish to reddish pattern

*Baliosus nervosus

26b    Middle of each elytron with 8 rows of punctures … 27

27a    Elytra entirely dark and pronotum mostly red

Anisostena ariadne
*Chalepus bicolor

27b    Elytra and pronotum of same colors

Elytra uniformly black to dark metallic bluish

Anisostena nigrita

Elytra light or dark brownish with variable blackish, brownish or reddish patterns

*Sumitrosis inaequalis
*Sumitrosis rosea

28a (from 22) Third tarsal segment not deeply lobed [HELP] … Subfamily Chrysomelinae … 29

28b    Third tarsal segment deeply lobed [HELP] … Subfamily Eumolpinae (large part) … 34

29a    Dorsum dark without color pattern

Punctures of elytron confused, not ranged in rows

*Gastrophysa cyanea
Plagiodera versicolora

Punctures of elytron arranged in rows

Chrysolina cribraria
Phaedon desotonis
Phaedon viridis
*Phratora americana

29b    Dorsum with color pattern … 30

30a    Elytra dark with narrow reddish border

Microtheca ochroloma

30b    Elytra not as above … 31

31a    Elytra with one or more complete stripes

A. Low flattened species, small and parallel-sided

*Prasocuris vittata

AA. High convex species, larger and rounded

Larger beetle, length more than 7 mm, dark stripes black

*Leptinotarsa decemlineata
*Leptinotarsa juncta

Smaller beetle, length less than 6.5 mm, dark stripes brown

*Calligrapha bidenticola
*Zygogramma suturalis

31b    Elytral with spots or irregular pattern … 32

32a    Pronotum with sides longitudinally impressed or grooved [HELP]

Elytral color orange, red or pinkish with a pattern of irregular black bands

*Chrysomela interrupta
*Chrysomela knabi

Elytral color yellowish with broken darker stripes

*Chrysomela scripta

32b    Sides of pronotum not impressed … 33

33a    Legs black

Labidomera clivicollis

33b    Legs pale

*Calligrapha ignota
Calligrapha multipunctata
Calligrapha philadelphica
*Calligrapha pnirsa
Calligrapha rowena
Calligrapha scalaris
Calligrapha spiraeae

34a (from 28) Middle and hind tibia with distal outer margin excavated and usually with a stiff brush of setae [HELP] … 35

34b    Middle and hind tibia not modified as above … 37

35a    Larger beetle, length greater than 6.5mm, metallic green or bluish in color

*Typophorus nigritus viridicyaneus

35b    Smaller beetle, not metallic … 36

36a    Side of prothorax with strongly expanded lobe that is directed forward and partially covers head and mouthparts [HELP]

*Paria fragariae
Paria opacicollis
Paria quadriguttata
*Paria quadrinotata
*Paria scutellaris
Paria sellata
Paria sexnotata
*Paria thoracica

36b    Side of prothorax without forward-directed lobe that partially covers head and mouthparts [HELP]

Metachroma laevicolle
Metachroma laterale
Metachroma luridum
Metachroma marginale
Metachroma orientale
Metachroma pellucidum
Metachroma quercatum

37a (from 34) Dorsum hairy or scaly

Glyptoscelis barbata
*Glyptoscelis pubescens

37b    Dorsum not hairy or scaly … 38

38a    Large metallic green species, length greater than 10 mm, elytra smooth and polished without large punctures

*Chrysochus auratus

38b    Smaller species, length less than 4 mm, if metallic green then elytra with deep coarse punctures … 39

39a    Antennae with distal segments distinctly shorter and thicker than preceding segments; body robust and compact, dark shiny metallic

Side of prothorax with expanded lobe that is directed forward and partially covers head and mouthparts [HELP]

*Spintherophyta globosa

Side of prothorax without forward-directed lobe that partially covers head and mouthparts [HELP]

*Brachypnoea clypealis
*Brachypnoea convexa
*Brachypnoea margaretae
*Brachypnoea puncticollis
*Brachypnoea tristis

39b    Antennae with distal segments only slightly shorter and thicker than preceding segments, body less robust, may or may not be metallic … 40a


40a    Elytra of one dark color, without raised longitudinal ridges and rows of punctures

Rhabdopterus picipes
*Rhabdopterus praetextus
*Tymnes chrysis
Tymnes metasternalis
*Tymnes tricolor

40b    Elytra brownish or brownish overlain with greenish reflection, with raised longitudinal ridges and rows of punctures

*Colaspis brunnea
*Colaspis pini
Colaspis suilla

41a (from 23) Hind leg with femur enlarged for jumping [HELP] … Subfamily Galerucinae, Tribe Alticini … 42

41b    Hind leg with femur narrow, not modified for jumping … 65

42a    Claw segment of hind tarsus greatly swollen … 43

42b    Claw segment of hind tarsus normal … 46

43a    Punctures of elytra arranged in rows … 44

43b    Punctures of elytra confused, not formed into rows … 45

44a    Dorsum hairy

Distigmoptera apicalis
Distigmoptera pilosa

44b    Dorsum without hairs

Pachyonychus paradoxus

45a (from 43) Pronotum narrow, as long as broad

Pachyonyhchis paradoxus

45b    Pronotum broad, distinctly wider than long

A. Body generally smaller, depressed; margins of elytra and pronotum more strongly explanate

Capraita circumdata
Capraita pervittata
Capraita scalaris
*Capraita sexmaculata
*Capraita subvittata
Capraita thyamoides

AA. Body generally larger, more robust; margins of elytra and pronotum less expanded

B. Elytra with stripes

Kuschelina fallax
Kuschelina preplexa
Kuschelina petaurista
Kuschelina weismani

BB. Elytral without stripes

Elytra black

Kuschelina concinna
*Kuschelina vians

Elytra pale with slightly darker sutural area

*Kuschelina suturella

Elytra entirely metallic blue or green

*Kuschelina gibbitarsa

Elytra metallic blue or green with pale margin under humeri

Kuschelina thoracica

46a (from 42) Antenna with only 10 segments

Psylliodes convexior
Psylliodes punctulatus

46b    Antenna with 11 segments … 47

47a    Spur of hind tibia broad at apex and bifurcate [HELP]

*Dibolia borealis
Dibolia chelones

47b    Spur of hind tibia narrow at apex, with a single point … 48

48a    Hind tibia with angular dilation or tooth on outer margin before apex [HELP]… 49

48b    Hind tibia with outer margin before apex without modification [HELP] … 50

49a    Large species, length greater than 4.0 mm; elytron pale with irregular brownish pattern

*Blepharida rhois

49b    Small species, length less than 2.5 mm; elytra of one uniform color

A. Hind tibia with series of irregular teeth above angular tooth; body shiny black, not metallic [HELP]

Pseudodibolia opima

AA. Hind tibia without series of teeth above angular tooth; body shiny or dull, usually with distinct metallic reflection [HELP]

B. Head with median area of vertex with distinct punctures [HELP]

Punctures of elytral base confused

Chaetocnema irregularis

Punctures of elytral base in regular rows

*Chaetocnema denticulata
*Chaetocnema minuta
Chaetocnema pinquis
*Chaetocnema producta

BB. Head with median area of vertex without punctures

Chaetocnema alutacea
*Chaetocnema confinis
Chaetocnema crenulata
*Chaetocnema pulicaria
*Chaetocnema truncata

50a (from 48) Pronotum before base with lateral furrow on each side and/or distinct transverse impression [HELP] … 51

50b    Pronotum before base without lateral furrows or transverse impression … 58

51a    Elytra with punctures in rows … 52

51b    Elytra with punctures confused … 57

52a    Hind margin of pronotum weakly sinuate on each side [HELP] … 53

52b    Hind margin of pronotum straight … 56

53a    Dorsal surface conspicuously hairy

Color black or nearly black

*Epitrix brevis
*Epitrix cucumeris
*Epitrix fuscula
*Epitrix lobatus

Color brownish

*Epitrix hirtipennis
Epitrix humeralis

53b    Dorsal surface without conspicuous hairs … 54

54a    Pronotum without transverse impression before base (only longitudinal furrow on each side)

Mantura floridana

54b    Pronotum with transverse impression before base in addition to short longitudinal furrow on each side

Pronotum red, elytral blackish

Hornaltica bicolor

Pronotum and elytra of one color

*Acallepitrix nitens
*Margaridisa atriventris

55a    Lateral margin of pronotum with series of small teeth

*Orthaltica copalina


55b    Lateral margin of pronotum without series of small teeth … 56

56a    Pronotum red, elytra metallic

*Derocrepis erythropus
Trichaltica scabricula

56b    Pronotum matching color of elytra

Dorsal color brownish overlain with greenish metallic reflection

*Crepidodera longula
Derocrepis aesculi

Dorsal coloration entirely metallic

*Crepidodera browni
*Crepidodera nana
Crepidodera vaga
*Crepidodera violacea
*Derocrepis carinata

57a (from 51) Pronotum metallic

Altica aeneola
*Altica chalybea
Altica ignita
*Altica kalmiae
Altica litigata
Altica vicaria
Altica woodsi

57b    Pronotum yellow to orange

Elytral yellow to orange, same color as pronotum

*Parchicola tibialis

Elytra metallic blue

*Parchicola iris

58a (from 50) Punctures of elytra arranged in rows

Cerataltica insolita
Glyptina spuria

58b    Punctures of elytra confused … 59

59a    First segment of hind tarsus at least half as long as hind tibia

Color or dorsum very dark brown to blackish

Longitarsus melanurus

Color of dorsum medium brown to yellowish or reddish brown

*Longitarsus pratensis
Longitarsus suspectus
Longitarsus testaceus

59b    First segment of hind tarsus less than half as long as tibia … 60

60a    Small species, length less than 3.0 mm

A. Elytra metallic, pronotum reddish

*Lupraea picta

AA. Elytral black with pale markings, pronotum black

Pale markings of elytra are spots

Phyllotreta bipustulata
Phyllotreta conjuncta
Phyllotreta striolata
(small part)

Pale markings of elytra are stripes

Phyllotreta liebecki
*Phyllotreta striolata
(large part)
*Phyllotreta zimmermanni

60b    Large species, length greater than 3.0 mm … 61

61a    Body elongate and depressed … 62

61b    Body broad and robust … 63

62a    Hind femur weakly enlarged, face with strong groove above eye [HELP]

Elytra brownish or blackish without metallic reflection

*Luperaltica nigripalpis

Elytra dark with metallic reflection

Luperaltica senilis

62b    Hind femur strongly enlarged, face without groove above eye [HELP]

Elytra entirely black

*Systena frontalis
*Systena hudsonias

Elytra yellowish without pale stripe

*Systena marginalis

Elytra blackish to yellowish, each with pale stripe

*Systena blanda
*Systena elongata

63a (from 61) Pronotum and elytra of uniform red color

*Strabala rufa

63b    Pronotum and elytra not of one uniform color … 64

64a    Elytra with stripes

Extreme lateral margins of elytra black

*Disonycha glabrata

Extreme lateral margins of elytra pale

Disonycha alternata
Disonycha arizonae
Disonycha caroliniana
Disoncyha pennsylvanica
Disonycha pluriligata
Disonycha uniguttata
Disonycha weismani

64b    Elytra without stripes

A. Elytra orange-red with large, common black spot

*Disonycha discoidea (large part)

AA. Elytra uniformly dark

Pronotum pale without dark spots

Disonycha collata
*Disonycha xanthomelas

Pronotum pale with small dark spots

Disonycha triangularis

65a (from 41) Dorsal surface conspicuously hairy … Subfamily Galerucinae, Tribe Galerucini (large part) … 66

65b    Dorsal surface without conspicuous hairs … 68

66a    Large species, length greater than 6.7 mm; elytra with uniform well-defined stripes

Derospidea brevicollis
Trirhabda canadensis
*Trirhabda virgata

66b    Smaller species, length less than 6.5 mm, with or without stripes … 67


67a    Elytra brownish with black to brownish stripes that may be distinct and complete ranging to faint or abbreviated

Ophraella americana
Ophraella communa
Ophraella conferta
Ophraella notata
Ophraella notulata
Ophraella sexvittata

67b    Elytra not as above

A. Pronotum with dark spots

Pronotum with three dark spots; elytra yellowish brown, each with an incomplete dark submarginal stripe and dark basal spot

*Xanthogaleruca luteola

Pronotum with impressed dark spot on each side; elytra dark with a narrow pale border or dark with vague pale median stripe

Galerucella nymphaeae

AA. Pronotum without dark spots

Elytra pale brownish without stripes, densely hairy and finely punctate

Tricholochmaea tuberculata

Elytra reddish without stripes, sparsely hairy and coarsely punctate

Tricholochmaea cavicollis
Tricholochmaea rufosanquinea

68a (from 65) Larger species, length greater than 10 mm

Monocesta coryli

68b    Smaller species, length less than 7 mm … 69

69a    Elytra uniformly of one dark color, usually metallic bluish

Pronotum and elytra dark metallic

Scelolyperus bimarginatus
Scelolyperus cyanellus
*Scelolyperus liriophilus
*Scelolyperus meracus

Pronotum entirely reddish or black with a large reddish spot

*Phyllecthris dorsalis

69b    Elytra not uniformly dark … 70

70a    Tarsal claws bifid [HELP] … 71

70b    Tarsal claws each with broadly angular tooth [HELP] … 73

71a    Elytra surface without longitudinal impressions or raised ridges

Elytra yellowish to greenish with round black spots; punctures fine

*Diabrotica undecimpunctata howardi

Elytra yellowish brown without round black spots; punctures coarse

Ophraella cribrata

71b    Elytra surface with longitudinal impressions and raised ridges … 72

72a    Elytra black and yellow striped

Acalymma vinctum
*Acalymma vittatum

72b    Elytra without distinct stripes

Diabrotica barberi
*Diabrotica virgifera virgifera

73a (from 70) Elytron pale with a very broad dark median stripe, body subparallel-sided

Pronotum entirely pale

Phyllobrotica antennata
Phyllobrotica circumdata
*Phyllobrotica limbata
Phyllobrotica vittata

Pronotum pale with dark lateral areas

Phyllecthris gentilis

73b    Elytron yellow or reddish, usually with a fragmented black pattern or sometimes nearly entirely pale; body robust, sides rounded

*Ceratoma trifurcata



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Thursday, February 12, 2004