Tentative Chechlist of the Polistine Tribe Mischocyttarini
by
James M. Carpenter (American Museum of Natural
History)
Tribe Mischocyttarini Carpenter, 1993
Genus Mischocyttarus
de Saussure
Mischocyttarus de Saussure, 1853, Ét. Fam. Vesp. 2: 19, genus (2 species).
Type species: Zethus labiatus
Fabricius, 1804, by subsequent designation of Ashmead, 1902, Can. Ent. 34: 166.
Mischocytharus de Saussure, 1853, Ét. Fam. Vesp. 1,
Introduction: viii, footnote. Incorrect
original spelling of Mischocyttarus
de Saussure.
Myschocytharus de Saussure, 1853, Ét. Fam. Vesp. 2: 3,
footnote. Incorrect spelling of Mischocyttarus de Saussure.
Miscocyttarus de Saussure, 1854, Ét. Fam. Vesp. 2: 256. Incorrect
spelling of Mischocyttarus de
Saussure.
Mischocittarus de Saussure, 1857, Ét. Fam. Vesp. 2: xi. Incorrect
spelling of Mischocyttarus de
Saussure.
Myschocyttarus Schrottky, 1904, Allg. Zeitschr. Ent. 9: 345. Incorrect spelling of Mischocyttarus de Saussure.
Mischocythorus Krombein, 1979, in
Krombein et al., Cat. Hym. Am. N. Mex. 2: 1516. Incorrect spelling of Mischocytharus de Saussure.
Subgenus
Clypeopolybia Brèthes
Clypeopolybia Brèthes, 1923, Rev. Fac. Nac. Agron. Univ. La Plata 15: 16, genus.
Type species: Clypeopolybia duckei
Brèthes, 1923 [=Polistes flavicans Fabricius, 1804], by monotypy.
adjectus Zikán
Polybia mexicana ?; Fox, 1898,
Proc. Acad. Nat. Sci. Philad.: 451 [partim].
- Bequaert, 1933, Entomologica Am. 13: 142 (probably M. alfkenii).
Mischocyttarus
adjectus Zikán, 1935, Arq. Inst. Biol. Veget. 1
(3): 154 (key), 188, figs. 56, 59, female - [
Distr.: Brazil: Rio de Janeiro.
carbonarius
carbonarius (de Saussure)
Polybia carbonaria de Saussure, 1854, Ét. Fam. Vesp. 2: 198, female
(in division My) - “Le Brésil”
(London?); 1855, Ét. Fam. Vesp. 2.: pl. XXVI fig. 5. - Dalla Torre, 1894, Cat.
Hym. 9: 162 (cat.).
Megacanthopus carbonarius; Ducke, 1905,
Bol. Mus. Goeldi, Pará 4: 690; 1907, Bol. Mus. Para. Emilo Goeldi 5: 190.
Megacanthopus
flavicans var. carbonaria;
Ducke, 1910, Ann. Hist.-Nat.
Mus. Natl. Hung. 8: 539.
Mischocyttarus flavicans carbonarius; Ducke, 1918, Rev. Mus.
Paul. 10: 351.
Mischocyttarus carbonarius; Bequaert,
1933, Entomologica Am. 13: 114 (compared to M.
flavitarsis). - Richards, 1941, Proc. R. Ent. Soc. London (B) 10: 125 (in
subgenus Mischocyttarus, flavicans group). - Bequaert, 1944,
Bull. Mus. Comp. Zool. 94: 261 (list; probable syn.: M. ruficornis Cameron). - Richards, 1945, Trans. R. Ent. Soc.
London 95: 311 (key), 336, figs. 23-24, pl. 4 figs. 181-182 (in subgenus Mischocyttarus, flavicans group). - Zikán, 1949, Bol. Parq. Nac. Itatiaia 1: 36
(key), 188, figs. 125, 274-275. - Silveira, 1998, Pap. Avuls. Zool., São Paulo
40 (23): 366 (morphology).
Mischocyttarus
carbonarius race
flavonotatus Zikán, 1949, Bol. Parq. Nac. Itatiaia 1: 36 (key), 189, female
- “Yungas de la Paz, (Bolivia)” (São Paulo). - Richards, 1978, Soc. Wasps Am.:
287 (syn. of M. carbonarius (de
Saussure)).
Mischocyttarus carbonarius carbonarius;
Richards, 1978, Soc. Wasps Am.: 283 (key), 287 (in subgenus Clypeopolybia). - Raw, 1989, Maracá
Rainfor. Proj., Invert. Limnol., Prelim. Rep.: 29 (resemblance to Stelopolybia angulata;
Distr.:
carbonarius tibialis Richards
Mischocyttarus
carbonarius tibialis Richards, 1978, Soc. Wasps
Am.: 283 (key), 287, female (in subgenus Clypeopolybia)
- “
Distr.: Costa Rica*; Colombia: Amazonas, Cauca, Nariño.
clypeatus Zikán
Mischocyttarus
clypeatus Zikán, 1935, Arq. Inst. Biol. Veget. 1 (3): 154
(key), 192, figs. 63-65, pl. 2 fig. 4, female, male, nest - [Brazil] “Itatiaya”
(lectotype female Inst. Oswaldo Cruz, Rio de Janeiro); 1949, Bol. Parq. Nac. Itatiaia
1: 24 (key), 122, figs. 66-67, 206-207, 326. - Richards, 1978, Soc. Wasps Am.:
282 (key), 284 (designation of lectotype; in subgenus Clypeopolybia).
Distr.: Brazil: Rio de Janeiro.
duckei
(du Buysson)
Megacanthopus Duckei du Buysson, 1908, Rev. Ent., Caen 27: 218, female
- “Guyane française; Nouveau Chantier du Bas Maroni” (Paris). - Carpenter,
1999, Am. Mus. Novitat. 3259: 20 (notes on type).
Mischocyttarus duckei; Richards, 1940, Ann. Mag. Nat. Hist. (11) 5: 183
(possibly same as M. rufidens (de Saussure));
1941, Proc. R. Ent. Soc. London (B) 10: 125 (in
subgenus Mischocyttarus, flavicans group). - Bequaert, 1944,
Bull. Mus. Comp. Zool. 94: 261 (list). - Richards, 1945, Trans. R. Ent. Soc.
London 95: 311 (key), 337 (in subgenus Mischocyttarus,
flavicans group). - Zikán, 1949, Bol.
Parq. Nac. Itatiaia 1: 36 (key), 187. - Richards, 1978, Soc. Wasps Am.: 283
(key), 285 (in subgenus Clypeopolybia).
- Overal, 1978, Bol. Mus. Para. Emílio Goeldi (N. S.), Zool. 94: 10 (list). -
Almeida and Castro, 1991, Acta Biol. Par., Curitiba 20: 49 (coll. list).
Distr.: Guyana; French Guiana; Brazil:
Amazonas.
flavicans coroico Richards
Mischocyttarus
flavicans var. coroico
Richards, 1945, Trans. R. Ent. Soc. London 95: 311 (key), 335, fig. 21,
female (in subgenus Mischocyttarus, flavicans group) - “BOLIVIA: Coroico”
(Berlin); also from Chaco; 1978, Soc. Wasps Am.: 283 (key), 288 (in subgenus Clypeopolybia).
Distr.: Bolivia: La Paz.
flavicans flavicans (Fabricius)
Polistes flavicans Fabricius, 1804, Syst. Piez.: 276 - “America
meridionali” (Kobenhavn).
Polybia testacea; de Saussure, 1854, Ét. Fam. Vesp. 2: 183, male
not female (syn.: Polistes flavicans
Fabricius) [misidentification]. - Zikán, 1949, Bol. Parq. Nac. Itatiaia 1: 211
(original description composite).
Polybia flavicans; Dalla Torre, 1894, Cat. Hym. 9: 163 (syn.: Polistes testacea Fabricius; cat.);
1904, Gen. Ins. 19: 77 (cat.).
Megacanthopus goeldii Ducke, 1905, Bol. Mus. Goeldi
4: 690, female, male, nest - [Brazil] “Barcellos nos igapós do rio Negro”
(Bern, Buenos Aires); 1908, Deutsch. Ent. Z. 1908: 699 (Brazil: Pará); 1918,
Rev. Mus. Paul. 10: 350 (syn. of M.
flavicans (Fabricius)). - Nascimento and Overal, 1979, Bol. Mus. Para.
Emílio Goeldi (N. S.) 95: 9 (list). - Carpenter, 1999, Am. Mus. Novitat. 3259: 21 (notes on types).
Mischocyttarus flavicans; Ducke, 1918,
Rev. Mus. Paul. 10: 350. - Bequaert, 1944, Bull. Mus. Comp. Zool. 94: 261
(list). - Richards, 1941, Proc. R. Ent. Soc. London (B) 10: 125 (in subgenus Mischocyttarus, flavicans group), 128 (compared to heliconius n. sp.); 1945, Trans. R. Ent. Soc. London 95: 311 (key),
333, figs. 19-20, 25-26 (in subgenus Mischocyttarus,
flavicans group; syn.: Megacanthopus goeldii Ducke). - Zikán,
1949, Bol. Parq. Nac. Itatiaia 1: 36 (key), 186, figs. 124, 273. - Overal,
1978, Bol. Mus. Para. Emílio Goeldi (N. S.), Zool. 94: 10 (list). - Silveira,
1998, Pap. Avuls. Zool., São Paulo 40 (23): 364-366, fig. 8b (compared to M. aripuanaensis n. sp.).
Clypeopolybia
Duckei Brèthes, 1923, Rev. Fac. Nac. Agron. Univ.
La Plata 15: 16, female, male, pl. III fig. 3 - “Bolivia”
(Buenos Aires) [junior secondary homonym of M.
duckei (du Buysson, 1908)]. - Zikán, 1949, Bol. Parq. Nac. Itatiaia 1: 231.
- Richards, 1978, Soc. Wasps Am.: 287 (syn. of M. flavicans (Fabricius)).
Mischocyttarus
flavicans var. flavicans; Richards, 1945, Trans. R. Ent. Soc.
London 95: 311 (key), 335 (in subgenus Mischocyttarus,
flavicans group).
Mischocyttarus flavicans flavicans; Richards, 1978, Soc. Wasps
Am.: 283 (key), 288 (in subgenus Clypeopolybia).
Distr.:
heliconius Richards
Mischocyttarus
heliconius Richards, 1941, Proc. R. Ent. Soc.
London (B) 10: 125 (key), 128, female (in subgenus Kappa, heliconius group)
- “BRITISH GUIANA: Essequibo R., Moraballi Creek, high forest” (
Mischocyttarus
heliconius var. heliconius; Richards, 1945, Trans. R. Ent. Soc.
London 95: 311 (key), 339 (in subgenus Kappa,
heliconius group).
Mischocyttarus
heliconius var. iperuae
Richards, 1945, Trans. R. Ent. Soc. London 95: 311 (key), 340, female (in
subgenus Kappa, heliconius group) - “British Guiana, Essequibo R., Moraballi Creek,
Wallaba (Iperua spp., mimoseae) Bush”
(
Mischocyttarus
heliconius morph heliconius; Richards, 1978, Soc. Wasps Am.: 282 (key), 284 (in
subgenus Clypeopolybia).
Mischocyttarus
heliconius morph iperuae; Richards, 1978, Soc. Wasps Am.: 282 (key), 284 (in
subgenus Clypeopolybia).
Distr.:
piger Richards
Mischocyttarus
piger Richards, 1945, Trans. R. Ent. Soc. London
95: 311 (key), 340, fig. 28, female (in subgenus Kappa, heliconius group)
- “
Distr.:
richardsi Zikán
Mischocyttarus
richardsi Zikán, 1949, Bol. Parq. Nac.
Itatiaia 1: 24 (key), 123, figs. 68-69, 208-209, 371, female, male - [Brazil]
“Nova Teutônia, Sul de Santa Catarina” (lectotype female Inst. Oswaldo Cruz,
Distr.:
sericeus Richards
Mischocyttarus
sericeus Richards, 1978, Soc. Wasps Am.: 282 (key),
283, female (in subgenus Clypeopolybia)
- “
Distr.: Bolivia: La Paz.
wygodzinskyi Zikán
Mischocyttarus
wygodzinskyi Zikán, 1949, Bol. Parq. Nac. Itatiaia 1: 24 (key), 120, figs.
204, 292, female - [Brazil] “Rio de Janeiro, D. F.” (Inst. Oswaldo Cruz, Rio de
Janeiro). - Richards, 1978, Soc. Wasps Am.: 282 (key), 285 (in subgenus Clypeopolybia).
Distr.: Brazil: Rio de Janeiro, São Paulo.
woytkowskyi Richards
Mischocyttarus
woytkowskyi Richards, 1978, Soc. Wasps Am.: 283
(key), 286, male (in subgenus Clypeopolybia)
- “PERU: Cuzco, Valle Cosñipata, Hacienda Maria, 400 m” (Tucumán).
Distr.: Peru: Cuzco.
Subgenus
Haplometrobius Richards
Haplometrobius Richards, 1978, Soc. Wasps Am.: vi, 17, 274, 389, subgenus of Mischocyttarus de Saussure.
Type species: Mischocyttarus iheringi
Zikàn, 1935, by original designation.
alboniger Richards
Mischocyttarus
alboniger Richards, 1978, Soc. Wasps Am.: 392
(key), 411, female (in subgenus Haplometrobius)
- “SURINAM: Nassau Mts” (Leiden). - Raw, 1989, Maracá Rainfor. Proj., Invert.
Limnol., Prelim. Rep.: 29 (nest; Brazil), 36 (distr.); 1989, Maracá Rainfor.
Proj., Invert. Limnol., Prelim. Rep.: 41 (habitat preference). - Sarmiento,
1994, Rev. Biol. Trop. 42 (1/2): 361 (alboniguer
[!]; Colombia: Amazonas, Caquetá, Vaupés; list). - Kojima and van Achterberg,
1997, Zool. Med., Leiden 71: 159 (list).
Distr.: Colombia (?): Amazonas, Caquetá,
Vaupés; Suriname; Brazil: Roraima.
annulatus Richards
Mischocyttarus
annulatus Richards, 1978, Soc. Wasps Am.: 392
(key), 411, female (in subgenus Haplometrobius)
- “BRAZIL: Mato Grosso, base camp, 12°50’S, 31°47’W” (London).
Distr.: Brazil: Mato Grosso.
artifex Ducke
Mischocyttarus artifex Ducke, 1914, Zool. Jahrb.
(Syst.) 36: 315, 327, fig. J, female, male - [Brazil] “Obidos
am unteren Amazonas” (lectotype female São Paulo); 1918, Rev. Mus. Paul. 10:
354. - da Fonseca, 1926, Rev. Mus. Paul. 14: 183, pl. 2 figs. 1-2 (nest). -
Bequaert, 1933, Entomologica Am. 13: 118 (nest). - Richards, 1941, Proc. R.
Ent. Soc. London (B) 10: 128 (in subgenus Megacanthopus,
artifex group); 1945, Trans. R. Ent.
Soc. London 95: 321 (key), 446, figs. 105-106, 115 (in subgenus Megacanthopus, artifex group). - Zikán, 1949, Bol. Parq. Nac. Itatiaia 1: 22
(key), 102, figs. 59, 192, 306, 346. - Richards, 1978, Soc. Wasps Am.: 395
(key), 426 (designation of lectotype, as Megacanthopus
[!]; in subgenus Haplometrobius).
- Overal, 1978, Bol. Mus. Para. Emílio Goeldi (N. S.), Zool. 94: 10
(list). - Nascimento and Overal, 1979, Bol. Mus. Para. Emílio Goeldi (N. S.)
95: 10 (list). - Sarmiento, 1994, Rev. Biol. Trop. 42 (1/2): 362 (Colombia;
list). - Cooper, 1998, Ent. Mon. Mag. 134: 293, 296 (key), 305 (in artifex group; additional description;
Brazil; Colombia), fig. 6. - Carpenter, 1999, Am. Mus. Novitat. 3259: 20 (notes
on type).
Distr.: Colombia: Amazonas, Putumayo, Amazonas; Brazil:
Pará.
bahiaensis Zikán
Mischocyttarus
bahiaënsis Zikán, 1949, Bol. Parq. Nac. Itatiaia 1: 20 (key), 88, female -
[Brazil] “Estado da Bahia” (São Paulo). - Richards, 1978, Soc. Wasps Am.: 390 (key), 401 (in subgenus Haplometrobius). - Carpenter, 1999, Am. Mus. Novitat. 3259: 20
(notes on type).
Distr.: Brazil: Bahia, São Paulo.
bequaertii Richards
Mischocyttarus
bequaertii Richards, 1945, Trans. R. Ent. Soc.
London 95: 320 (key), 428, fig. 91, male, female (in subgenus Megacanthopus, prominulus group) - “COLOMBIA: Dept. Meta, Restrepo, 500 m.”
(holotype male Cambridge); 1978, Soc. Wasps Am.: 392 (key), 411 (in subgenus Haplometrobius). - Sarmiento, 1994, Rev.
Biol. Trop. 42 (1/2): 361 (list).
Distr.: Colombia: Meta.
capichaba Zikán
Mischocyttarus
indeterminabilis; Zikán, 1935, Arq. Inst. Biol.
Veget. 1 (3): 154 (key), 168 (compared to paraguayensis
n. sp. and iheringi n. sp.).
Mischocyttarus
capichaba Zikán, 1949, Bol. Parq. Nac. Itatiaia 1:
19 (key), 83, fig. 43, female - “Estado do Espírito Santo, Brasil” (São Paulo).
- Richards, 1978, Soc. Wasps Am.: 393 (key), 416 (in subgenus Haplometrobius). - Cooper, 1998, Ent.
Mon. Mag. 134: 293 (characters).
Distr.:
cerberus acheron Richards
Mischocyttarus
cerberus var. acheron
Richards, 1940, Ann. Mag. Nat. Hist. (11) 5: 179, female - “British Guiana:
Mazaruni Settlement” (
Mischocyttarus cerberus acheron;
Richards, 1978, Soc. Wasps Am.: 395 (key), 425 (in
subgenus Haplometrobius). -
Sarmiento, 1994, Rev. Biol. Trop. 42 (1/2): 362 (list).
Distr.:
cerberus cerberus Ducke
Polybia pallipes; Fox, 1898,
Proc. Acad. Nat. Sci. Philad.: 450 [partim]. Misidentification.
Megacanthopus cerberus “Buyss. n. sp.”
Ducke, 1910, Rev. Ent.,
Mischocyttarus
cerberus Ducke, 1918, Rev. Mus. Paul. 10: 371 (probably var. of M. surinamensis (de Saussure)) -
[Brazil] “Maranhão: Caxias. Connu du bas Amazon” (lectotype male London). -
Richards, 1940, Ann. Mag. Nat. Hist. (11) 5: 178
(treated as species described by Ducke, 1918); 1941, Proc. R. Ent. Soc. London
(B) 10: 128 (in subgenus Megacanthopus,
cerberus group). - Bequaert, 1944,
Bull. Mus. Comp. Zool. 94: 261 (typical form). - Richards, 1945, Trans. R. Ent.
Soc. London 95: 442, figs. 103-104.- Zikán, 1949, Bol. Parq. Nac. Itatiaia 1:
96. - Overal, 1978, Bol. Mus. Para. Emílio Goeldi (N. S.), Zool. 94: 10 (list).
- Richards, 1978, Soc. Wasps Am.: 424 (designation of lectotype, from Pará). -
Nascimento and Overal, 1979, Bol. Mus. Para. Emílio Goeldi (N. S.) 95: 9 (list). - Carpenter and van der Vecht,
1991, Ann. Carnegie Mus. 60 (3): 217 (note on material
seen by Fox). - Silveira, 2004, Rev. Bras. Entomol. 48 (3): 335 (in cerberus
group), 338 (key).
Mischocyttarus
cerberus var. cerberus;
Richards, 1940, Ann. Mag. Nat. Hist. (11) 5: 179 (female, male;
Mischocyttarus
cerberus cerberus;
Zikán, 1949, Bol. Parq. Nac. Itatiaia 1: 21 (key). - Richards, 1978, Soc. Wasps
Am.: 395 (key), 425 (in subgenus Haplometrobius).
- Sarmiento, 1994, Rev. Biol. Trop. 42 (1/2): 362 (
Distr.: Colombia (?): Caquetá, Putumayo; Suriname;
Brazil: Maranhão, Mato Grosso, Pará.
cerberus styx Richards
Mischocyttarus
cerberus var. styx
Richards, 1940, Ann. Mag. Nat. Hist. (11) 5: 180, female - “Brazil: São Paulo, Dumont” (London); 1945, Trans. R.
Ent. Soc. London 95: 321 (key), 445, pl. 2 figs. 150-151 (in subgenus Megacanthopus, cerberus group).
Mischocyttarus souzalopesi Zikán, 1949, Bol. Parq. Nac. Itatiaia 1:
85, female not male (Brazil: Goiás).
Misidentification.
Mischocyttarus varolii Zikán, 1949, Bol. Parq.
Nac. Itatiaia 1: 21 (key), 95, figs. 52-53, 187-188, 361, male, female -
[Brazil] “Araçatuba” (lectotype male Inst. Oswaldo Cruz, Rio de Janeiro); also
from two other localities in São Paulo. - Richards, 1978, Soc. Wasps Am.: 425
(designation of lectotype; syn. of M.
cerberus styx Richards).
Mischocyttarus
cerberus styx;
Richards, 1978, Soc. Wasps Am.: 395 (key), 425 (syn.: M. varolii Zikán; in subgenus Haplometrobius).
- Almeida and Castro, 1991, Acta Biol. Par., Curitiba 20: 49 (coll. list). -
Giannotti, 1998, Rev. Bras. Ent. 41 (2-4): 217-224 (ethology); 1999,
Sociobiology 33 (3): 325-338 (ethology).
Distr.: Brazil: Goiás, Mato Grosso, São Paulo.
chanchamayoensis Richards
Mischocyttarus “chanchamayoanus Zikán nov. spec.”
Weyrauch, 1943, Bol. Mus. Hist. Nat. Javier Prado 7: 68. Nomen
nudum.
Mischocyttarus
chanchamayoensis Richards, 1978, Soc. Wasps Am.: 393 (key), 414, female (in
subgenus Haplometrobius) - “PERU:
Junín, Valle Chanchamayo, 800 m” (Tucumán). - Cooper, 1998, Ent. Mon. Mag. 134:
91 (in iheringi group; male, nest;
Colombia; Ecuador; Peru: Pasco). - Colomo and Berta, 2005, Rev. Soc. Entomol.
Argent. 64: 78 (list).
Distr.: Colombia: Caqueta, Putumayo; Ecuador:
Morona-Santiago, Napo; Peru: Junín, Pasco.
chloroecus Cooper
Mischocyttarus
chloroecus Cooper, 1998, Ent. Mon. Mag. 134: 295 (key), 300, fig. 10, male,
female, nest (in subgenus Haplometrobius,
artifex species group) - “COLOMBIA:
Putumayo, Mocoa, 550m” (holotype male London); also from Caquetá; and Ecuador.
Distr.: Colombia: Putumayo, Caquetá; Ecuador:
Morona-Santiago.
claretianus Zikán
Mischocyttarus
claretianus Zikán, 1949, Bol. Parq. Nac. Itatiaia 1: 21 (key), 101, figs.
58, 409, female - [Brazil] “Fonte Ahu, Curitiba, Estado do Paraná” (Inst.
Oswaldo Cruz, Rio de Janeiro). - Richards, 1978, Soc. Wasps
Am.: 396 (key), 431 (in subgenus Haplometrobius).
Distr.: Brazil: Paraná.
cleomenes Richards
Mischocyttarus
cleomenes Richards, 1945, Trans. R. Ent. Soc.
London 95: 320 (key), 432, figs. 93, 95, pl. 1 fig. 121, female, male, nest (in
subgenus Megacanthopus, surinamensis group) - “MEXICO: N.
Yucatán, Temax” (holotype female London); also from Panama; 1978, Soc. Wasps
Am.: 391 (key), 406 (in subgenus Haplometrobius).
- West-Eberhard et al., 1995, Hym. Costa
Rica: 585 (Costa Rica). - Rodríguez-Palafox, 1996, in Llorente-Bousquets
et al., Biodiv. Taxon. Biogeogr. Artrópod. México: 482 (list).
Distr.: Mexico: Yucatán; Costa Rica; Panama.
confusoides Zikán
Mischocyttarus
confusoides Zikán, 1949, Bol. Parq. Nac. Itatiaia 1: 236,
sex and locality not stated (lectotype female, Inst. Oswaldo Cruz,
Distr.: Brazil: Rio de Janeiro.
confusus Zikán
Mischocyttarus
confusus Zikán, 1935, Arq. Inst. Biol. Veget. 1 (3): 154
(key), 195, figs. 66, 69, 73, pl. 3 fig. 10, female - [Brazil] “Itatiaya, 700
m.” (Tucumán). - Richards, 1941, Proc. R. Ent. Soc. London (B) 10: 127 (in
subgenus Megacanthopus, surinamensis group); 1945, Trans. R.
Ent. Soc. London 95: 320 (key), 433, fig. 96 (in subgenus Megacanthopus, surinamensis
group). - Zikán, 1949, Bol. Parq. Nac. Itatiaia 1: 21 (key), 99, 236 (“host” of
confusoides n. sp.), figs. 57, 191,
367. - Richards, 1978, Soc. Wasps Am.: 391 (key), 404 (in subgenus Haplometrobius). - Colomo and Berta,
2005, Rev. Soc. Entomol.
Argent. 64: 78 (note on type).
Distr.:
Brazil: Rio de Janeiro.
cooperi Richards
Mischocyttarus
cooperi Richards, 1978, Soc. Wasps Am.: 418,
female, male (in subgenus Haplometrobius)
- “
Distr.:
curitybanus Zikán
Mischocyttarus
curitybanus Zikán, 1949, Bol. Parq. Nac. Itatiaia
1: 22 (key), 106, female - [
Distr.: Brazil: Rio de Janeiro, Paraná.
decimus Richards
Mischocyttarus
decimus Richards, 1978, Soc. Wasps Am.: 390 (key),
400, female (in subgenus Haplometrobius)
- “
Distr.: Peru: Loreto; Brazil: Pará; Bolivia: Beni.
dimorphus Zikán
Mischocyttarus “dimorphus Zikán nov. spec.” Weyrauch, 1943, Bol. Mus. Hist. Nat.
Javier Prado 7: 69. Nomen nudum.
Mischocyttarus dimorphus Zikán, 1949, Bol. Parq. Nac. Itatiaia 1: 19 (key), 84, fig. 44, male, female - “Valle
Chanchamayo, 1.200 m, Peru” (lectotype male Inst. Oswaldo Cruz, Rio de
Janeiro). - Richards, 1978, Soc. Wasps Am.: 396 (key), 432 (designation of
lectotype; in subgenus Haplometrobius).
- Cooper, 1998, Ent. Mon. Mag. 134: 293, 295 (key; in artifex group). - Silveira, 2004, Rev. Bras. Entomol. 48 (3): 336
(in cerberus group), 338 (key). - Colomo and Berta, 2005, Rev. Soc.
Entomol. Argent. 64: 79 (note on types).
Distr.: Peru: Junín.
elegantulus Zikán
Mischocyttarus
“elegantulus Zikán
nov. spec.” Weyrauch, 1943,
Bol. Mus. Hist. Nat. Javier Prado 7: 68.
Nomen nudum.
Mischocyttarus elegantulus Zikán, 1949, Bol. Parq. Nac. Itatiaia 1:
23 (key), 114, fig. 200, female - “Tingo Maria, Rio Harallaga [Huallaga], 700
m, (Peru)” (Inst. Oswaldo Cruz, Rio de Janeiro). - Richards, 1978, Soc. Wasps Am.: 392 (key), 410 (in subgenus Haplometrobius). - Sarmiento, 1994, Rev. Biol. Trop. 42 (1/2): 361
(Colombia: Putumayo; list).
Distr.: Colombia: Putumayo; Ecuador: Napo (?); Peru:
Huánuco.
filipendulus Cooper
Mischocyttarus
filipendulus Cooper, 1998, Ent. Mon. Mag. 134: 295, 296 (key), 301
(mimicry), 302, fig. 7, female, nest (in subgenus Haplometrobius, artifex
species group) - “COLOMBIA: Amazonas, La Chorrera” (London); also from
Putumayo.
Distr.: Colombia: Putumayo, Amazonas.
garbei Zikán
Mischocyttarus
garbei Zikán, 1935, Arq. Inst. Biol. Veget. 1 (3): 153
(key), 194, female - [
Mischocyttarus
garbei var. garbei;
Richards, 1945, Trans. R. Ent. Soc. London 95: 320 (key), 438 (in subgenus Megacanthopus, surinamensis group).
Mischocyttarus
flaveolus Zikán, 1949, Bol. Parq. Nac. Itatiaia 1:
20 (key), 89, female - [Brazil] “Estado do Espírito Santo” (São Paulo). -
Richards, 1978, Soc. Wasps Am.: 404 (syn. of M. garbei Zikán).
Distr.: Brazil: Espírito Santo, Rio de Janeiro.
giffordi Raw
Mischocyttarus
giffordi Raw, 1985, Rev. Bras. Ent. 29: 107, figs. 1-8, female, male -
[Brazil] “Olho de água da Onça, Fazenda água Limpa...Brasília, D. F.” (holotype female Rio de Janeiro). -
Cooper, 1998, Ent. Mon. Mag. 134: 293 (characters).
Distr.:
Brazil: D. F.
hoffmanni
Zikán
Mischocyttarus hoffmanni Zikán, 1949, Bol. Parq. Nac. Itatiaia 1: 19
(key), 86, figs. 46, 183, female - [Brazil] “Rio Laeisse, próximo a Nova
Bremen, Estado de Santa Catarina” (lectotype Inst. Oswaldo
Cruz, Rio de Janeiro); also from Rio Grande do Sul and São Paulo. - Richards,
1978, Soc. Wasps Am.: 390 (key), 403 (designation of lectotype; in subgenus Haplometrobius).
Distr.: Brazil: Santa Catarina, São Paulo.
ignotus Zikan
Mischocyttarus
ignotus Zikán, 1949, Bol. Parq. Nac. Itatiaia 1: 20 (key), 91, fig. 49,
female - “Franca, Estado S. Paulo, Brasil” (Inst. Oswaldo
Cruz, Rio de Janeiro). - Richards, 1978, Soc. Wasps Am.: 390 (key), 403 (in
subgenus Haplometrobius). -
Sarmiento, 1994, Rev. Biol. Trop. 42 (1/2): 361 (Colombia: Valle; list).
Distr.: Colombia (?): Meta, Putumayo, Valle; Brazil: Rio
Grande do Sul, Rio de Janeiro (?), São Paulo.
iheringi Zikán
Mischocyttarus
iheringi Zikán, 1935, Arq. Inst. Biol. Veget. 1 (3): 154
(key), 177, figs. 41, 43, 47, pl. 1 fig. 6, female, male, nest - [Brazil]
“Südhang des Itatiaya, ...700 bis 1100 m ü. d. M.” (lectotype male Inst.
Oswaldo Cruz, Rio de Janeiro). - Richards, 1941, Proc. R. Ent. Soc. London (B)
10: 127 (in subgenus Megacanthopus, iheringi group); 1945, Trans. R. Ent.
Soc. London 95: 321 (key), 440, fig. 102 (in subgenus Megacanthopus, artifex
group). - W. Zikán and Wydodzinsky, 1948, Bol. Serv. Nac. Pesq. Agron. 4: 74
(list). - Zikán, 1949, Bol. Parq. Nac. Itatiaia 1: 22 (key), 107, 236 (“host”
of iheringoides n. sp.), figs. 60-61,
363-364. - Richards, 1978, Soc. Wasps Am.: 394 (key), 421 (designation of
lectotype; syn. M. iheringoides Zikán;
in subgenus Haplometrobius). -
Cooper, 1998, Ent. Mon. Mag. 134: 293 (characters). - Colomo and Berta, 2005,
Rev. Soc. Entomol. Argent. 64: 80 (note on types).
Mischocyttarus
iheringoides Zikán, 1949, Bol. Parq. Nac. Itatiaia
1: 236, locality not stated - (lectotype male Inst. Oswaldo Cruz,
Distr.: Brazil: Rio de Janeiro.
illusorius Richards
Mischocyttarus
illusorius Richards, 1978, Soc. Wasps Am.: 396 (key), 432, female (in
subgenus Haplometrobius) - “PERU:
Cuzco, Rio Urubamba, Santa Ana, 1100 m” (London). - Sarmiento, 1994, Rev. Biol.
Trop. 42 (1/2): 362 (Colombia: Nariño, Putumayo; list). -
Cooper, 1998, Ent. Mon. Mag. 134: 293, 295 (key; in artifex group). - Silveira, 2004, Rev. Bras. Entomol. 48 (3): 336
(in cerberus group), 338 (key).
Distr.: Colombia: Nariño, Putumayo;
Peru: Cuzco.
interjectus Zikán
Mischocyttarus
interjectus Zikán, 1935, Arq. Inst. Biol. Veget. 1
(3): 154 (key), 175, figs. 39-40, pl. 3 fig. 4, female, nest - [Brazil] “Gelände
der Fazenda ‘Valparaizo’ (etwa 550 m. Seehöhe)” (Inst. Oswaldo Cruz, Rio de
Janeiro); 1949, Bol. Parq. Nac. Itatiaia 1: 18 (key), 80, 236 (“host” of interjectoides n. sp.), figs. 42, 180,
365. - Richards, 1978, Soc. Wasps Am.: 396 (key), 433 (syn.: M. interjectoides Zikán; in subgenus Haplometrobius).
Mischocyttarus
interjectoides Zikán, 1949, Bol. Parq. Nac.
Itatiaia 1: 236, sex and locality not stated (lectotype male Inst. Oswaldo
Cruz,
Distr.: Brazil: Rio de Janeiro.
interruptus Richards
Mischocyttarus
interruptus Richards, 1978, Soc. Wasps Am.: 395 (key), 426, female (in
subgenus Haplometrobius) - “BRAZIL:
probably Amazonas, Rio Autaz” (Stockholm). - Sarmiento, 1994, Rev. Biol. Trop.
42 (1/2): 362 (Colombia; list). - Cooper, 1998, Ent. Mon. Mag. 134: 295 (key),
301 (mimicry), 304 (in artifex group;
variation; male; nest; Colombia; Peru).
Distr.: Colombia: Amazona; Peru: Loreto; Brazil:
Amazonas.
lemoulti (du Buysson)
Megacanthopus Le Moulti du Buysson, 1908, Rev. Ent., Caen 27: 217, male -
“Guyane française: Nouveau chantier du Bas Maroni” (Paris).
Mischocyttarus lemoulti; Bequaert, 1944,
Bull. Mus. Comp. Zool. 94: 262 (list). - Richards, 1978, Soc. Wasps Am.: 390
(key), 398 (in subgenus Haplometrobius).
Distr.:
leucoecus Cooper
Mischocyttarus
leucoecus Cooper, 1998, Ent. Mon. Mag. 134: 293,
296 (key), 301, fig. 4, male, female, nest (in subgenus Haplometrobius, artifex
species group) - “
Distr.: Colombia: Putumayo; Ecuador: Pastaza.
longicornis Zikán
Mischocyttarus “longicornis Zikán nov. spec.” Weyrauch, 1943, Bol. Mus. Hist. Nat.
Javier Prado 7: 68. Nomen nudum.
Mischocyttarus longicornis Zikán, 1949, Bol. Parq. Nac. Itatiaia 1: 22 (key), 110, figs. 63, 195-196, 384, female, male -
“Oxapampa, 1.700 m, Peru” (lectotype male Inst. Oswaldo Cruz, Rio de Janeiro).
- Richards, 1978, Soc. Wasps Am.: 393 (key), 419 (designation of lectotype; in
subgenus Haplometrobius).
Distr.: Peru: Pasco.
macarenae Cooper
Mischocyttarus
macarenae Cooper, 1998, Ent. Mon. Mag. 134: 90,
female, nest (in subgenus Haplometrobius,
iheringi group) - “COLOMBIA: Meta, La
Macarena” (London).
Distr.: Colombia: Meta.
maculipennis Cooper
Mischocyttarus
maculipennis Cooper, 1998, Ent. Mon. Mag. 134: 295,
296 (key), 303, female (in subgenus Haplometrobius,
artifex species group) - “COLOMBIA:
Putumayo, Villa Garzón 400m” (London).
Distr.:
melanoleucus Richards
Mischocyttarus
melanoleucus Richards, 1978, Soc. Wasps Am.: 392
(key), 410, female (in subgenus Haplometrobius)
- “
Distr.:
melanoxanthus Richards
Mischocyttarus
melanoxanthus Richards, 1978, Soc. Wasps Am.: 391
(key), 405, fig. 118, female (in subgenus Haplometrobius)
- “BRAZIL: Mato Grosso, camp site 12°50’S, 51°47’W” (London). - Sarmiento,
1994, Rev. Biol. Trop. 42 (1/2): 361 (
Distr.:
mendax Richards
Mischocyttarus
mendax Richards, 1978, Soc. Wasps Am.: 390 (key),
398, female (in subgenus Haplometrobius)
- “FRENCH GUIANA: Alicot-Oyapock” (
Distr.:
minifoveatus Cooper
Mischocyttarus
minifoveatus Cooper, 1998, Ent. Mon. Mag. 134: 89,
female, nest (in subgenus Haplometrobius)
- “PERU: Junin, Quebrada Mala Noche c. 10 km east of San Ramon, 1,000-1,300 m”
(London); 1998, Ent. Mon. Mag. 134: 301 (mimicry).
Distr.:
mirificus Zikán
Mischocyttarus
mirificus Zikán, 1935, Arq. Inst. Biol. Veget. 1
(3): 154 (key), 169, figs. 29, 31, 36, 38, pl. 2 fig. 1, female, male, nest - [
Distr.: Colombia: Putumayo; Peru: Loreto; Brazil: Rio de
Janeiro.
montivagus Cooper
Mischocyttarus
montivagus Cooper, 1996, Ent. Mon. Mag. 132: 273 (key), 277, fig. 4,
female, male, nest (in subgenus Haplometrobius,
mendax species group) - “PERU: Pasco,
Villa Rica, 1,500 m” (holotype female London); also from Bolivia.
Distr.: Peru: Pasco; Bolivia: La Paz.
moronae Cooper
Mischocyttarus
moronae Cooper, 1996, Ent. Mon. Mag. 132: 273 (key), 277, figs. 10-11, female
(in subgenus Haplometrobius, mendax species group) - “ECUADOR:
Morona-Santiago, Cordillera de Cutucu 6 km E. of Macas, 1,100 m” (London); also
from Tungurahua.
Distr.: Ecuador: Morona-Santiago,
Tungurahua.
narinensis Cooper
Mischocyttarus
narinensis Cooper, 1998, Ent. Mon. Mag. 134: 293,
295 (key), 297, figs. 1-2, 8, male, female, nest (in subgenus Haplometrobius, artifex species group) - “COLOMBIA: Nariño, Barbacoas (50m)”
(holotype male London). - Silveira, 2004, Rev. Bras. Entomol. 48 (3): 336 (in cerberus
group), 338 (key).
Distr.: Colombia: Nariño.
naumanni Richards
Mischocyttarus
naumanni Richards, 1978, Soc. Wasps Am.: 393 (key),
418, female, male (in subgenus Haplometrobius)
- “ECUADOR: Napo, Limoncocha, 00°24’S, 76°36’W” (holotype female London). -
Cooper, 1998, Ent. Mon. Mag. 134: 90 (note on male), 92 (in iheringi group; nest; Colombia), fig. 1.
Distr.: Colombia: Caqueta, Putumayo; Ecuador: Napo.
nigroclavatus Zikán
Mischocyttarus
nigroclavatus Zikán, 1949, Bol. Parq. Nac. Itatiaia 1: 20 (key), 92, fig.
50, male, female - “San José da Costa Rica” (lectotype male Inst. Oswaldo Cruz, Rio de Janeiro). - Richards, 1978, Soc. Wasps Am.: 397
(key), 435 (designation of lectotype; in subgenus Haplometrobius). - West-Eberhard et al., 1995, Hym.
Distr.: Costa Rica; Panama; Colombia: Nariño.
nigropygialis Zikán
Mischocyttarus “nigropygialis Zikán nov. spec.” Weyrauch, 1943, Bol. Mus. Hist. Nat.
Javier Prado 7: 68. Nomen nudum.
Mischocyttarus nigropygialis Zikán, 1949, Bol. Parq. Nac. Itatiaia 1: 23 (key), 111, figs. 197, 368, female - “Oxapampa, 1.300
m, Peru” (Inst. Oswaldo Cruz, Rio de Janeiro). - Richards, 1978, Soc. Wasps
Am.: 394 (key), 420 (in subgenus Haplometrobius).
- Cooper, 1998, Ent. Mon. Mag. 134: 301 (nigropygidialis
[!]; mimicry).
Distr.:
nomurae Richards
Mischocyttarus
nomurae Richards, 1978, Soc. Wasps Am.: 396 (key),
430, female, nest (in subgenus Haplometrobius)
- “
Distr.:
occultus Cooper
Mischocyttarus
occultus Cooper, 1996, Ent. Mon. Mag. 132: 274
(key), 277, figs. 3, 9, female, male, larva, nest (in subgenus Haplometrobius, mendax species group) - “
Distr.:
oecothrix Richards
Mischocyttarus
oecothrix Richards, 1940, Ann. Mag. Nat. Hist. (11)
5: 180, female, male - “British Guiana:
Distr.: Colombia: Nariño; Guyana; Peru: Junín (?)*;
Brazil: Amazonas*.
ornatus
Zikán
Mischocyttarus “ornatus Zikán
nov. spec.” Weyrauch, 1943, Bol. Mus. Hist. Nat. Javier Prado 7: 68. Nomen
nudum.
Mischocyttarus ornatus Zikán, 1949, Bol. Parq. Nac. Itatiaia 1: 22 (key),
109, figs. 62, 193-194, 366, male, female - “Oreja de Capello, 1.600 m., Peru”
(lectotype male Inst. Oswaldo Cruz, Rio de Janeiro). - Richards, 1978, Soc. Wasps Am.: 394 (key), 421 (designation of lectotype; in subgenus Haplometrobius). - Sarmiento, 1994, Rev.
Biol. Trop. 42 (1/2): 362 (Colombia: Caquetá; list). - Cooper, 1998, Ent. Mon.
Mag. 134: 91-92 (in iheringi group;
compared to chanchamayoensis
Richards; nest; Peru; Bolivia);
1998, Ent. Mon. Mag. 134: 300 (note on nest).
Distr.: Colombia: Caquetá; Peru: Junín; Bolivia: La Paz.
pallidus Zikán
Mischocyttarus
surinamensis; Zikán, 1935, Arq. Inst. Biol. Veget. 1 (3): 154 (key), 193.
Mischocyttarus
pallidus Zikán, 1949, Bol. Parq. Nac. Itatiaia 1: 23 (key), 116, fig. 202,
female - [Brazil] “Manaus” (São Paulo). - Richards, 1978, Soc. Wasps Am.: 390 (key), 402 (in subgenus Haplometrobius).
Distr.: Peru: Loreto (?); Brazil:
Amazonas.
paulistanus Zikán
Mischocyttarus
paulistanus Zikán, 1935, Arq. Inst. Biol. Veget. 1 (3): 154 (key), 176, female -
[Brazil] “Villa Emma, Staat S. Paulo” (São Paulo); 1949, Bol. Parq. Nac. Itatiaia 1: 21 (key), 99, fig. 56. - Richards, 1978, Soc.
Wasps Am.: 391 (key), 407 (designation of lectotype; in subgenus Haplometrobius). - Carpenter, 1999, Am.
Mus. Novitat. 3259: 22 (note
on type).
Mischocyttarus confusus; Richards, 1945, Trans. R. Ent. Soc. London
95: 434 [partim]. Misidentification.
Distr.: Brazil: Rio de Janeiro, São Paulo.
peruanus Zikán
Mischocyttarus “peruanus Zikán nov. spec.” Weyrauch, 1943, Bol. Mus. Hist. Nat.
Javier Prado 7: 68. Nomen nudum.
Mischocyttarus peruanus Zikán, 1949, Bol. Parq. Nac. Itatiaia 1: 20 (key),
94, figs. 51, 186, male - “Santa Ana, 1.100 m, Peru” (Inst. Oswaldo Cruz, Rio
de Janeiro). - Richards, 1978, Soc. Wasps Am.: 391 (key), 406 (in subgenus Haplometrobius). - Silveira, 2004, Rev.
Bras. Entomol. 48 (3): 336 (in cerberus group),
338 (key; probable syn. of M. cerberus).
Distr.: Peru: Cuzco.
peruviensis Richards
Mischocyttarus
peruviensis Richards, 1945, Trans. R. Ent. Soc.
London 95: 321 (key), 451, figs. 114, 117, female (in subgenus Megacanthopus, artifex group) - “PERU: Col. Perene, El Campamiento” (Cambridge);
also from Bolivia: Santa Cruz; 1978, Soc. Wasps Am.: 396 (key), 429 (in subgenus
Haplometrobius). - Cooper, 1998, Ent.
Mon. Mag. 134: 295 (key), 301 (mimicry), 304 (in artifex group; additional description; male; nest; Colombia;
Ecuador; Peru; Bolivia), fig. 11.
Distr.: Colombia: Putumayo, Amazonas; Ecuador: Morona-
Santiago; Peru: Junín, Pasco; Bolivia: Cochabamba, Santa Cruz.
polymorphus Cooper
Mischocyttarus
polymorphus Cooper, 1998, Ent. Mon. Mag. 134: 296 (key), 299, figs. 3, 5,
9, male, female, nest (in subgenus Haplometrobius,
artifex species group) - “COLOMBIA:
Putumayo, Mocoa (500m)” (holotype male London); also from two other localities
in Putumayo; and Caquetá; Amazonas.
Distr.: Colombia: Putumayo, Caquetá, Amazonas.
prominulus Richards
Mischocyttarus
prominulus Richards, 1941, Proc. R. Ent. Soc. London (B) 10: 127 (key),
129, female, male (in subgenus Megacanthopus,
prominulus group) - “BRITISH GUIANA:
Mazaruni R., Kartabo” (holotype female London); also from Bolivia. - Bequaert,
1944, Bull. Mus. Comp. Zool. 94: 263 (list). - Richards, 1945,
Trans. R. Ent. Soc. London 95: 320 (key), 425, figs. 89, 92, pl. 1 fig. 133 (in
subgenus Megacanthopus, prominulus group). - Zikán, 1949, Bol.
Parq. Nac. Itatiaia 1: 22 (key), 104. - Richards, 1978, Soc. Wasps Am.: 392
(key), 412, fig. 120 (in subgenus Haplometrobius).
- Raw, 1989, Maracá Rainfor. Proj., Invert. Limnol., Prelim. Rep.: 30 (Brazil),
37 (distr.); 1989, Maracá Rainfor. Proj., Invert. Limnol., Prelim. Rep.: 41
(habitat preference).
Mischocyttarus
prominulus var. prominulus; Richards, 1945, Trans. R. Ent. Soc.
London 95: 321 (key).
Distr.: Guyana; French Guiana*; Ecuador:
Napo; Brazil: Amazonas, Pará, Roraima; Bolivia.
reclusus Cooper
Mischocyttarus
reclusus Cooper, 1996, Ent. Mon. Mag. 132: 274
(key), 278, figs. 7, 14, female, larva, nest (in subgenus Haplometrobius, mendax
species group) - “COLOMBIA: Caqueta, Yurayacu (73 km S.W. of Florencia),
350-500 m” (London); also from Putumayo.
Distr.: Colombia: Caqueta, Putumayo.
reflexicollis Zikán
Mischocyttarus “reflexicollis Zikán nov. spec.” Weyrauch, 1943, Bol. Mus. Hist. Nat.
Javier Prado 7: 69. Nomen nudum.
Mischocyttarus reflexicollis Zikán, 1949, Bol. Parq. Nac. Itatiaia 1:
22 (key), 104, fig. 362, female - “Tingo Maria, 700 m, Rio Huallaga, Peru”
(lectotype Inst. Oswaldo Cruz, Rio de Janeiro). - Richards, 1978, Soc. Wasps Am.: 393 (key), 416 (designation of lectotype; in subgenus Haplometrobius). - Sarmiento, 1994, Rev.
Biol. Trop. 42 (1/2): 362 (Colombia: Amazonas, Putumayo; list). - Cooper, 1998,
Ent. Mon. Mag. 134: 293, 296 (key), 306 (in artifex
group; Colombia; Ecuador; Bolivia).
Distr.: Colombia: Amazonas, Putumayo;
Ecuador: Morona-Santiago; Peru: Huánuco; Bolivia: Cochabamba.
saussurei Zikán
Mischocyttarus
saussurei Zikán, 1949, Bol. Parq. Nac. Itatiaia 1:
21 (key), 98, figs. 54-55, 189-190, female, male - [Brazil] “Ipiranga, São
Paulo” (holotype female São Paulo). - Richards, 1978, Soc. Wasps Am.: 393
(key), 417 (in subgenus Haplometrobius).
- Sarmiento, 1994, Rev. Biol. Trop. 42 (1/2): 362 (Colombia: Caquetá; list). -
Cooper, 1998, Ent. Mon. Mag. 134: 90 (male identified by Richards as saussurei may be macarenae n. sp.).
Distr.: Brazil: São Paulo.
schadei Zikán
Mischocyttarus
schadei Zikán, 1949, Bol. Parq. Nac. Itatiaia 1: 17 (key), 70, figs. 30-31,
303, male - “Hohenau, Alto Paraná, Paraguai” (Inst. Oswaldo Cruz, Rio de
Janeiro). - Richards, 1978, Soc. Wasps Am.: 395 (key), 423 (in subgenus Haplometrobius). - Garcete-Barrett, 1999, Avispas Soc.
Parag.: 25 (key, fig. 9a), 27. - Silveira, 2004, Rev.
Bras. Entomol. 48 (3): 336 (belongs to artifex group).
Distr.: Brazil: Minas Gerais (?); Paraguay: Itapúa.
silvicola Zikán
Mischocyttarus “silvicola Zikán nov. spec.” Weyrauch, 1943, Bol. Mus. Hist. Nat.
Javier Prado 7: 68. Nomen nudum.
Mischocyttarus silvicola Zikán, 1949, Bol. Parq. Nac. Itatiaia 1: 23 (key),
115, figs. 65, 201, 290, female, male - “Valle do Chanchamayo, 800 e 1.200 m,
(Peru)” (lectotype male Inst. Oswaldo Cruz, Rio de Janeiro). - Richards, 1978,
Soc. Wasps Am.: 392 (key), 409, fig. 119 (designation
of lectotype; in subgenus Haplometrobius).
- Sarmiento, 1994, Rev. Biol. Trop. 42 (1/2): 361 (
Distr.:
souzalopesi Zikán
Mischocyttarus
souza-lopesi Zikán, 1949, Bol. Parq. Nac. Itatiaia
1: 19 (key), 85, figs. 45, 182, male not female - “Ilha Sêca, Estado de S.
Paulo (Brasil)” (lectotype male Inst. Oswaldo Cruz,
Distr.:
surinamensis occidentalis Richards
Mischocyttarus
surinamensis var. occidentalis Richards, 1945, Trans. R. Ent. Soc. London 95: 320
(key), 435 (in subgenus Megacanthopus,
surinamensis group) - “BRITISH
GUIANA: R. Potaro, Kaieteur Plateau, Tukeit trail, high forest” (holotype
female
Mischocyttarus surinamensis occidentalis; Richards, 1978, Soc. Wasps
Am.: 390 (key), 402 (in subgenus Haplometrobius).
Distr.:
Trinidad; Guyana.
surinamensis
surinamensis (de
Saussure)
Polybia surinamensis de Saussure, 1854, Ét. Fam.
Vesp. 2: 200, female, male (in division My)
- “
Megacanthopus surinamensis; Ducke,
1904, Bol. Mus. Goeldi, Pará 4: 360 (key), 361 (
Mischocyttarus surinamensis;
Ducke, 1918, Rev. Mus. Paul. 10: 354 [partim].
- da Fonseca, 1926, Rev. Mus. Paul. 14: 181 (compared to ypiranguensis n. sp.). - Vesey-Fitzgerald, 1939, Proc. R. Ent. Soc.
London (A) 14: 104-105 (mimicry). - Richards, 1940,
Ann. Mag. Nat. Hist. (11) 5: 182; 1941, Proc. R. Ent. Soc. London (B) 10: 127
(in subgenus Megacanthopus, surinamensis group). - Bequaert, 1944,
Bull. Mus. Comp. Zool. 94: 263 (list). - Richards, 1945, Trans. R. Ent. Soc.
London 95: 320 (key), 436, figs. 94, 97, 100, pl. 2 figs. 142-143 (in subgenus Megacanthopus, surinamensis group). - Zikán, 1949, Bol. Parq. Nac. Itatiaia 1: 20
(key), 89, figs. 48, 185, 289, 345. - Overal, 1978, Bol. Mus. Para. Emílio
Goeldi (N. S.), Zool. 94: 10 (list). - Raw, 1988, Entomologist 107: 106 (role
in agriculture).
Mischocyttarus
surinamensis var. surinamensis; Richards,
1945, Trans. R. Ent. Soc. London 95: 320 (key), 436 (in subgenus Megacanthopus, surinamensis group).
Mischocyttarus surinamensis surinamensis; Richards, 1978, Soc. Wasps
Am.: 390 (key), 401 (in subgenus Haplometrobius).
- Raw, 1989, Maracá Rainfor. Proj., Invert. Limnol., Prelim. Rep.: 37 (
Distr.: Colombia: Amazonas, Arauca,
Meta, Nariño, Vaupés; Guyana; Suriname; French Guiana; Ecuador: Napo; Peru:
Junín, Loreto; Brazil: Acre, Amazonas, Ceará, Maranhão, Mato Grosso, Pará, Rio
de Janeiro, Rondônia, Roraima; Bolivia (?).
sylvestris Richards
Mischocyttarus
sylvestris Richards, 1945, Trans. R. Ent. Soc.
London 95: 320 (key), 430, figs. 99, 101, pl. 1 fig. 120, female, male, nest
(in subgenus Megacanthopus, surinamensis group) - “BRITISH GUIANA:
Essequibo R., Moraballi Creek, dark forest” (holotype female London); 1978,
Soc. Wasps Am.: 390 (key), 397 (in subgenus Haplometrobius).
- Sarmiento, 1994, Rev. Biol. Trop. 42 (1/2): 361 (
Distr.:
synoecus Richards
Polybia
surinamensis;
Fox, 1898, Proc. Acad. Nat. Sci. Philad.: 448 [partim].
Mischocyttarus
synoecus Richards, 1940, Ann. Mag. Nat. Hist. (11)
5: 181, female, male - “British Guiana: Potaro R., Amatuk” (holotype female
Distr.:
tectus Cooper
Mischocyttarus
tectus Cooper, 1996, Ent. Mon. Mag. 132: 274, figs.
1, 8, 12-13, female, nest (in subgenus Haplometrobius,
mendax species group) - “
Distr.: Colombia: Putumayo; Peru: Huánuco.
tenuis
Richards
Mischocyttarus prominulus var. tenuis
Richards, 1945, Trans. R. Ent. Soc. London 95: 321
(key), 427, fig. 90, female (in subgenus Megacanthopus,
prominulus group) - “PERU: Dept.
Cuzco” (Berlin).
Mischocyttarus
tenuis; Richards, 1978, Soc. Wasps Am.: 392 (key),
413 (in subgenus Haplometrobius). -
Sarmiento, 1994, Rev. Biol. Trop. 42 (1/2): 361 (Colombia: Meta,
Putumayo; list).
Distr.: Colombia: Meta, Putumayo; Ecuador: Napo; Peru:
Cuzco, Huánuco; Brazil: Pará.
tertius Richards
Mischocyttarus
tertius Richards, 1978, Soc. Wasps Am.: 396 (key),
431, female (in subgenus Haplometrobius)
- “BRAZIL: Mato Grosso, base camp 12°50’S, 51°47’W” (London). - Cooper, 1998,
Ent. Mon. Mag. 134: 293 (characters).
Distr.:
thrypticus Richards
Mischocyttarus
thrypticus Richards, 1945, Trans. R. Ent. Soc.
London 95: 321 (key), 451, fig. 113, female, male (in subgenus Megacanthopus, artifex group) - “
Distr.:
Paraguay: Alto Paraná.
travassosi Zikán
Mischocyttarus travassosi Zikán, 1949, Bol. Parq. Nac.
Itatiaia 1: 19 (key), 82, fig. 181, female, male - [Brazil] “Rio de Janeiro, D.
F.” (lectotype male Inst. Oswaldo Cruz, Rio de Janeiro). - Richards, 1978, Soc.
Wasps Am.: 394 (key), 422 (designation of lectotype; in
subgenus Haplometrobius).
Distr.:
tricolor Richards
Mischocyttarus
garbei var. tricolor
Richards, 1945, Trans. R. Ent. Soc. London 95: 320 (key), 439, female (in
subgenus Megacanthopus, surinamensis group) - “
Mischocyttarus tricolor; Richards, 1978,
Soc. Wasps Am.: 390 (key), 400. - Garcete-Barrett, 1999, Avispas Soc. Parag.:
25 (key), 27 (? Paraguay).
Distr.: Brazil: Goiás, Mato Grosso, São Paulo; ? Paraguay:
Paraguarí, San Pedro, Canindeyú; Argentina: Misiones.
tunari Cooper
Mischocyttarus
tunari Cooper, 1996, Ent. Mon. Mag. 132: 274 (key), 279, fig. 5, female,
male (in subgenus Haplometrobius, mendax species group) - “BOLIVIA:
Cochabamba, Villa Tunari, 400 m” (holotype female London).
Distr.:
Bolivia: Cochabamba.
undulatus
(Ducke)
Megacanthopus undulatus Ducke, 1905, Rev. Ent. 24: 21, female, male -
[Brazil] “du poste brésilien sur l’Oyapoc (Etat du Pará)” (lectotype male
Belém); also from Amazonas (Teffé). - Overal, 1978, Bol. Mus. Para. Emílio Goeldi (N. S.), Zool. 94: 6 (punctatus [!]; designation of lectotype), 10 (list). - Nascimento
and Overal, 1979, Bol. Mus. Para. Emílio Goeldi (N. S.) 95: 10 (list). -
Carpenter, 1999, Am. Mus. Novitat. 3259: 23 (notes on type).
Distr.: Colombia: Putumayo; Brazil: Amapá, Amazonas, Mato
Grosso; Bolivia: Beni.
vredeni Richards
Mischocyttarus
vredeni Richards, 1978, Soc. Wasps Am.: 393 (key),
417, female, male (in subgenus Haplometrobius)
- “
Distr.:
weyrauchi Zikán
Mischocyttarus
“weyrauchi Zikán
nov. spec.” Weyrauch, 1943,
Bol. Mus. Hist. Nat. Javier Prado 7: 68.
Nomen nudum.
Mischocyttarus weyrauchi Zikán, 1949, Bol. Parq. Nac. Itatiaia 1: 23 (key),
113, figs. 64, 198-199, 369, female, male - “San Luiz, 800 m, Vale de
Chanchamayo, Peru” (lectotype male Inst. Oswaldo Cruz, Rio de Janeiro). -
Richards, 1978, Soc. Wasps Am.: 393 (key), 415
(designation of lectotype; in subgenus Haplometrobius).
- Sarmiento, 1994, Rev. Biol. Trop. 42 (1/2): 362 (
Distr.: Colombia: Putumayo; Ecuador:
Napo; Peru: Huánuco, Junín; Bolivia: Beni.
xanthocerus Richards
Mischocyttarus
garbei var. xanthocerus
Richards, 1945, Trans. R. Ent. Soc. London 95: 320 (key), 439, female, male
(in subgenus Megacanthopus, surinamensis group) - “Bolivia”
(holotype female Berlin). - Zikán, 1949, Bol. Parq. Nac. Itatiaia 1: 219.
Mischocyttarus
xanthocerus; Richards, 1978, Soc. Wasps Am.: 391
(key), 405 (in subgenus Haplometrobius).
- Sarmiento, 1994, Rev. Biol. Trop. 42 (1/2): 361 (list).
Distr.: Panama; Colombia: Nariño, Putumayo; Peru:
Huánuco; Bolivia: Beni.
ypiranguensis da Fonseca
Mischocyttarus
ypiranguensis da Fonseca, 1926, Rev. Mus. Paul. 14: 181, pl. 1 figs. 1-2,
male, female, nest - [Brazil] “Ypiranga” (lectotype male São Paulo). - Zikán,
1935, Arq. Inst. Biol. Veget. 1 (3): 154 (key), figs. 32, 34;
1949, Bol. Parq. Nac. Itatiaia 1: 17 (key), 69, figs. 28-29, 176, 285, 302,
340. - Richards, 1978, Soc. Wasps Am.: 395 (key), 427 (designation of
lectotype; in subgenus Haplometrobius).
- Cooper, 1998, Ent. Mon. Mag. 134: 293 (characters).. - Silveira, 2004, Rev.
Bras. Entomol. 48 (3): 336 (in artifex group).
Distr.: Brazil: São Paulo.
Subgenus
Kappa de Saussure
Kappa de Saussure, 1854, Ét. Fam. Vesp. 2: 200, division V of subgenus Polybia of genus Polybia Lepeletier (8 species); at pl. 25: division of genus Polybia.
Type species: Polybia injucunda de
Saussure, 1854, by subsequent designation of Bequaert, 1933, Entomologica Am.
13: 112.
Pseudopolybia H. von Ihering, 1896, Bull. Soc. Zool. Fr. 21: 159-162; Zool. Anz.
19: 452, genus (2 species). Junior
homonym of Pseudopolybia de Saussure,
1863.
Type species: “Polistes ignobilis Haliday, 1836 = Vespa
atra Olivier, 1791” [= Polybia socialis de Saussure, 1854] by subsequent designation of Bequaert,
1933, Entomologica Am. 13: 112.
acreanus Silveira
Mischocyttarus
acreanus Silveira, 2006, Zootaxa 1321: 4, 16 (key),
81, figs. 73, 75, female (in subgenus Kappa) - “Brazil, Acre, Rio Branco”
(Belem).
Distr.: Brazil: Acre.
adolphi Zikán
Megacanthopus
frontalis; Ducke, 1907, Bol. Mus. Para. Emilo
Goeldi 5: 189 [partim].
Mischocyttarus frontalis; Ducke, 1918, Rev. Mus. Paul. 10: 352 [partim].
Mischocyttarus duckei Zikán, 1935, Arq. Inst. Biol. Veget. 1 (3): 155
(key), 196, figs. 68, 70, 75, female - [Brazil] “Umgebung von Manáos (Rio
Negro, Staat Amazonas)” (Inst. Oswaldo Cruz, Rio de Janeiro). Junior secondary
homonym of M. duckei (du Buysson, 1908).
Mischocyttarus
adolphi Zikán, 1949, Bol. Parq. Nac. Itatiaia 1: 37
(key), 192 [replacement name for duckei
Zikán]. - Richards, 1978, Soc. Wasps Am.: 289 (key), 295 (in subgenus Kappa). - Silveira, 2006, Zootaxa 1321:
3 (male), 14 (key), 53, figs. 51-53, 65 (in subgenus Kappa; Brazil).
Distr.:
Brazil: Amapá, Amazonas, Pará.
anthracinus
Richards
Mischocyttarus frontalis var. anthracinus
Richards, 1945, Trans. R. Ent. Soc. London 95: 312 (key), 345, fig. 41,
female, male (in subgenus Kappa, ater group) - “PARAGUAY: Alto Parana,
Hohenau” (holotype female Cambridge).
Mischocyttarus anthracinus;
Zikán, 1949, Bol. Parq. Nac. Itatiaia 1: 37 (key), 193,
fig. 408. - Richards, 1978, Soc. Wasps Am.: 290 (key), 296 (in subgenus Kappa). - Garcete-Barrett, 1999, Avispas Soc.
Parag.: 26 (key, fig. 13a), 28. - Silveira, 2006,
Zootaxa 1321: 14 (key), 59, figs. 56, 62-63 (in subgenus Kappa).
Distr.:
bertonii Ducke
Polybia rufidens; Fox, 1898,
Proc. Acad. Nat. Sci. Philad.: 450 [partim].
Mischocyttarus
injucundus bertonii
Ducke, 1918, Rev. Mus. Paul. 10: 370, female, male - “
Mischocyttarus
metathoracicus var. picturatus Bequaert, 1938, Proc. R. Ent. Soc. London 7: 135 [partim].
Mischocyttarus
injucundus var. bertonii; Richards, 1945, Trans. R. Ent. Soc.
London 95: 313 (key), 351.
Mischocyttarus bertonii; Zikán, 1949,
Bol. Parq. Nac. Itatiaia 1: 38 (key), 194, fig. 129. - Richards, 1978, Soc.
Wasps Am.: 291 (key), 301 (in subgenus Kappa;
Distr.: Brazil: Mato Grosso do Sul, Rio Grande do Sul,
São Paulo; Bolivia: Santa Cruz; Paraguay: Central, Cordillera, Paraguarí,
Guairá, San Pedro, Concepción, Canindeyú, Alto Paraná.
deceptus (Fox)
Polybia
decepta Fox, 1895, Proc. California Acad. Sci. (2)
5: 269, female - locality not stated [
Mischocyttarus ater;
Ducke, 1918, Rev. Mus. Paul. 10: 351, 369 [partim].
Mischocyttarus socialis;
Bequaert, 1944 (1943), Ann. Ent. Soc. Am. 36: 721 [partim].
Mischocyttarus atrocyaneus Zikán, 1949, Bol. Parq. Nac.
Itatiaia 1: 40 (key), 204, fig. 404, female - “San José da Costa Rica” (Inst. Oswaldo Cruz, Rio de Janeiro). - Richards, 1978, Soc. Wasps Am.: 290
(key), 297 (in subgenus Kappa). -
Snelling, 1983, Pan-Pacif. Ent. 59: 275 (nest; Mexico; El Salvador; Panama). -
West-Eberhard et al., 1995, Hym.
Costa Rica: 585 (Costa Rica). - Rodríguez-Palafox, 1996, in
Llorente-Bousquets et al., Biodiv. Taxon. Biogeogr. Artrópod. México:
482 (list). - Silveira, 2006, Zootaxa 1321: 45 (syn. of M. deceptus
(Fox)).
Mischocyttarus
atramentarius; Richards, 1978, Soc. Wasps Am.: 298
[partim].
Mischocyttarus deceptus; Silveira, 2006, Zootaxa 1321: 4, 13 (key), 45 (syn.: M.
atrocyaneus Zikán; in subgenus Kappa; male;
Distr.: Mexico: Chiapas, Guerrero,
Jalisco, Nayarit, Oacaxa, Veracruz, Yucatán; Guatemala; El Salvador; Costa
Rica; Panama; Colombia; Santander; Ecuador (?).
efferus Silveira
Polybia
metathoracica; Fox, 1898, Proc. Acad. Nat. Sci. Philad.: 449 (
Mischocyttarus metathoracicus; Ducke,
1918, Rev. Mus. Paul. 10: 353 [partim].
Mischocyttarus
efferus Silveira, 2006, Zootaxa 1321: 4, 16 (key),
78, figs. 70-71, 76, 78, female, male, nest (in subgenus Kappa) - “Brazil,
Pará, I. do Marajó, P. de Pedras” (holotype female Belem); also from Maranhão;
Rio de Janeiro; and Bolivia; Paraguay.
Distr.:
fidus Silveira
Mischocyttarus
fidus Silveira, 2006, Zootaxa 1321: 3, 14 (key),
58, figs. 57-61, male (in subgenus Kappa)
- “
Distr.: Brazil: Pará.
fitzgeraldi Bequaert
Mischocyttarus
fitzgeraldi Bequaert, 1938, Proc. R. Ent. Soc.
Distr.:
flavoscutellatus Zikán
Mischocyttarus
flavoscutellatus Zikán, 1935, Arq. Inst. Biol.
Veget. 1 (3): 155 (key), 198, figs. 76-77, female, nest - [
Mischocyttarus metathoracicus var. flavoscutellatus; Richards, 1945, Trans.
R. Ent. Soc. London 95: 313 (key), 347, fig. 33 (in subgenus Kappa, ater group).
Mischocyttarus
flavoscutellatoides Zikán, 1949, Bol. Parq. Nac.
Itatiaia 1: 236, sex and locality not stated (lectotype female Inst. Oswaldo
Cruz,
Distr.: Brazil: Minas Gerais, Rio de
Janeiro, São Paulo.
frontalis (Fox)
Polybia
frontalis Fox, 1898, Proc. Acad. Nat. Sci. Philad.:
454, female, male - “
Megacanthopus frontalis;
Ducke, 1910, Ann. Hist.-Nat. Mus. Natl. Hung. 8: 540.
Mischocyttarus frontalis;
Ducke, 1918, Rev. Mus. Paul. 10: 352 [partim].
- Zikán, 1935, Arq. Inst. Biol. Veget. 1 (3): 155
(key). - Richards, 1941, Proc. R. Ent. Soc. London (B) 10: 125 (in subgenus Kappa, ater group); 1945, Trans. R. Ent. Soc. London 95: 312 (key), 344,
fig. 32 (in subgenus Kappa, ater group) [partim, not var.]. - Zikán, 1949, Bol. Parq. Nac. Itatiaia 1: 38
(key), 195. - Richards, 1978, Soc. Wasps Am.: 290 (key), 296 (in subgenus Kappa). - Overal, 1978, Bol. Mus. Para.
Emílio Goeldi (N. S.), Zool. 94: 10 (list). - Garcete-Barrett, 1999, Avispas Soc. Parag.: 26
(key), 28. - Silveira, 2006, Zootaxa 1321: 14 (key, fig. 13aa), 62 (in subgenus
Kappa; nest; Brazil; Paraguay).
Mischocyttarus frontalis
var. frontalis; Richards, 1945,
Trans. R. Ent. Soc. London 95: 312 (key), 345 (in subgenus Kappa, ater group)
Distr.:
Brazil: Goiás, Mato Grosso; Paraguay: San Pedro, Canindeyú.
funerulus
Zikán
Mischocyttarus funerulus Zikán, 1949, Bol. Parq.
Nac. Itatiaia 1: 38 (key), 197, 236 (“host” of funeruloides n. sp.), figs. 133, 402, female, male, nest - [
Mischocyttarus
funeruloides Zikán, 1949, Bol. Parq. Nac. Itatiaia
1: 236, sex and locality not stated - (lectotype female Inst. Oswaldo Cruz,
Distr.:
imitator (Ducke)
?
Vespa atra Olivier, 1792, Encyc. Méthod. 6 (2): 674 - [French Guiana:] “
Megacanthopus
imitator Ducke, 1904, Bol. Mus. Goeldi, Pará 4: 359
(key), 362, fig. a, pl. 1 fig. 5 (Polybia
imitatrix [!]), female, male, nest - [Brazil:] “Belem do Pará” (lectotype
female São Paulo); also from another locality; 1905, Bol. Mus. Goeldi, Pará 4:
689 (syn. of M. ater (Olivier,
1792)). - Richards, 1978, Soc. Wasps Am.: 296 (designation of lectotype). -
Nascimento and Overal, 1979, Bol. Mus. Para. Emílio Goeldi (N. S.) 95: 9
(list). - Carpenter, 1999, Am. Mus. Novitat. 3259: 21 (notes
on types).
Mischocyttarus ater var. uniformis Bequaert, 1938, Proc. R. Ent.
Soc.
Mischocyttarus socialis var. uniformis; Bequaert, 1944 (1943), Ann.
Ent. Soc. Am. 36: 722.
Mischocyttarus apicalis; Zikán, 1949,
Bol. Parq. Nac. Itatiaia 1: 40 (key), 204, figs. 139, 281. Misidentification.
Mischocyttarus
goyanus Zikán, 1949, Bol. Parq. Nac. Itatiaia 1: 40
(key), 206, female - “Estado de Goiás, Brasil” (Inst. Oswaldo Cruz, Rio de
Janeiro). - Richards, 1978, Soc. Wasps Am.: 290 (key), 297 (in subgenus Kappa). - Silveira, 2006, Zootaxa 1321:
49 (syn. of M. imitator Ducke).
Mischocyttarus imitator; Richards, 1978,
Soc. Wasps Am.: 290 (key), 296 (in subgenus Kappa).
- Silveira, 2006, Zootaxa 1321: 13 (key), 48, figs. 43-47, 97 (syns.: M. ater var. undiformis Bequaert; in subgenus Kappa; Bolivia;
Brazil; Peru; Venezuela).
Mischocyttarus
communalis Richards, 1978, Soc. Wasps Am.: 290
(key), 297, fig. 112, female (in subgenus Kappa)
- “
Distr.:
immarginatus Richards
? Mischocyttarus
rufidens; Zikán, 1935, Arq. Inst. Biol. Veget. 1 (3): 155 (key).
Mischocyttarus
immarginatus Richards, 1940, Ann. Mag. Nat. Hist.
(11) 5: 182, male, female - “
Mischocyttarus
immarginatoides Zikán, 1949, Bol. Parq. Nac.
Itatiaia 1: 236, figs. 352-353, locality not stated (holotype female Inst.
Oswaldo Cruz,
Distr.: Mexico: Chiapas, Colima, Guerrero,
Jalisco, Michoacán, Nayarit, Nuevo León, Oaxaca, Puebla, San Luis Potosí,
Sinaloa, Sonora (?), Tamaulipas, Veracruz, Yucatán, Zacatecas; Guatemala;
Honduras; El Salvador; Nicaragua; Costa Rica.
injucundus
(de Saussure)
Polybia injucunda de Saussure, 1854, Ét. Fam. Vesp. 2: 200, female
(in division Kappa)- [Brazil:] “Le
Para” (? lectotype Paris, syntypes Torino); 1855, Ét. Fam. Vesp. 2.: pl. XXV
fig. 8. - Smith, 1857, Cat. Hym. Ins. Coll. Brit. Mus.
5, Vesp.: 109. - Schulz, 1904 (1903), Sber. Akad. Bayer. Wiss. 33: 801 (male).
- Dalla Torre, 1904, Gen. Ins. 19: 77 (cat.). - von Ihering, 1904, Rev. Mus.
Paul. 6: 183. - Cameron, 1912, Timehri (3) 2: 208 (key), 211 (
Megacanthopus injucundus; Ducke, 1904,
Bol. Mus. Goeldi, Pará 4: 360 (key), 362; 1905, Bol. Mus. Goeldi, Pará
4: 689, pl. 4 fig. 22 (nest); 1907, Bol. Mus. Para. Emilo Goeldi 5: 189; 1910,
Ann. Hist.-Nat. Mus. Natl. Hung. 8: 540.
Polybia
bimarginata Cameron, 1912, Timehri (3) 2: 211,
female - locality not stated [
Mischocyttarus injucundus; Ducke, 1918,
Rev. Mus. Paul. 10: 353. - Zikán, 1935, Arq. Inst. Biol. Veget. 1 (3): 155
(key), figs. 67, 71. - Vesey-Fitzgerald, 1939, Proc. R. Ent. Soc. London (A)
14: 105 (mimicry). - Richards, 1941, Proc. R. Ent. Soc. London (B) 10: 125 (in
subgenus Kappa, ater group); 1945, Trans. R. Ent. Soc. London 95: 313 (key), 349,
figs. 35, 42, pl. 1 figs. 134-135, pl. 4 figs. 183-184 (in subgenus Kappa, ater group). - Bequaert, 1944, Bull. Mus. Comp. Zool. 94: 262
(typical form). - - Zikán, 1949,
Bol. Parq. Nac. Itatiaia 1: 39 (key), 201, figs. 135-136, 403. - Overal, 1978,
Bol. Mus. Para. Emílio Goeldi (N. S.), Zool. 94: 10 (list). - Silveira, 2006,
Zootaxa 1321: 17 (key), 92, figs. 5, 85, 87, 99-100 (syns.: M. injucundus
bimarginatus (Cameron), M. injucundus tingomariae Richards, M.
jucundus Richards, M. juruanus Richards; in subgenus Kappa; nest; Bolivia; Brazil; Colombia; Ecuador; Peru).
Mischocyttarus
injucundus bertonii;
da Fonseca, 1926, Rev. Mus. Paul. 14: 175.
Misidentification.
Mischocyttarus
“injucundus
nov. var. occidentalis Zikán”
Weyrauch, 1943, Bol. Mus. Hist. Nat. Javier Prado 7: 68. Nomen
nudum.
Mischocyttarus injucundus var. bimarginatus; Bequaert, 1944, Bull. Mus.
Comp. Zool. 94: 262. - Richards, 1945, Trans. R. Ent. Soc. London 95: 313
(key), 351, pl. 1 figs. 134-135, pl. 4, figs. 183-184 (in subgenus Kappa, ater group).
Mischocyttarus injucundus
var. injucundus; Richards, 1945,
Trans. R. Ent. Soc. London 95: 313 (key), 350 (in
subgenus Kappa, ater group)
Mischocyttarus bimarginatus; Zikán,
1949, Bol. Parq. Nac. Itatiaia
1: 214.
Mischocyttarus jucundus Richards, 1978, Soc. Wasps
Am.: 291 (key), 301, female (in subgenus Kappa)
- “
Mischocyttarus
juruanus Richards, 1978, Soc. Wasps Am.: 291 (key),
302, female (in subgenus Kappa) - “
Mischocyttarus injucundus
injucundus; Richards, 1978, Soc. Wasps Am.: 291 (key), 303 (in subgenus Kappa).
Mischocyttarus injucundus bimarginatus; Richards, 1978, Soc. Wasps
Am.: 291 (key), 303 (in subgenus Kappa).
- Raw, 1989, Maracá Rainfor. Proj., Invert. Limnol., Prelim. Rep.: 29 (nest;
resemblance to Polybia rejecta;
Mischocyttarus
injucundus tingomariae Richards, 1978, Soc. Wasps
Am.: 291 (key), 304, female, male (in subgenus Kappa) - “
Rhopalidia injucunda; Casolari and Casolari Moreno, 1979, Boll. Mus. Zool. Univ. Torino
(4): 97 (list: syntypes in Torino).
Distr.: Colombia: Amazonas, Putumayo; Ecuador: Napo;
Peru: Apurimac, Huánuco, Junín, Loreto, Ucayali; Venezuela; Trinidad; Guyana; French
Guiana*; Brazil: Amapá, Amazonas, Bahia (?), Maranhão, Pará, Roraima; Bolivia:
Beni.
latior (Fox)
Polybia
latior Fox, 1898, Proc. Acad. Nat. Sci. Philad.:
451, female - [
Mischocyttarus latior; Richards, 1941,
Proc. R. Ent. Soc. London (B) 10: 125 (in subgenus Kappa, ater group); 1945,
Trans. R. Ent. Soc. London 95: 312 (key), 344 (in subgenus Kappa, ater group; ?
syn.: M. villarricanus Zikán). -
Zikán, 1949, Bol. Parq. Nac. Itatiaia 1: 213. - Richards, 1978, Soc. Wasps Am.:
289 (key), 294 (in subgenus Kappa;
nest). - Almeida and Castro, 1991, Acta Biol. Par.,
Distr.:
latissimus Richards
Mischocyttarus
latissimus Richards, 1978, Soc. Wasps Am.: 289
(key), 294, female (in subgenus Kappa)
- “
Distr.: Colombia: Amazonas; Peru: Loreto; Bolivia: Beni,
Pando; Brazil: Amazonas.
laurae Silveira
Mischocyttarus
laurae Silveira, 2006, Zootaxa 1321: 3, 10 (key), 19, figs. 6-8, 11, 16,
female (in subgenus Kappa) - “Venezuela: Merida, Guarunes” (New York).
Distr.: Venezuela: Merida.
magdalensis Silveira
Mischocyttarus tolensis; Sarmiento, 1994, Rev.
Biol. Trop. 42 (1/2): 360 (Colombia: Magdalena, Nariño; list).
Mischocyttarus
magdalensis Silveira, 2006, Zootaxa 1321: 3, 11 (key), 29, figs. 27, 31,
female, nest (in subgenus Kappa) - “Colombia, Magdalena, N. Sierra
Nevada de Santa Marta, Rio Buritaca 100m” (London).
Distr.: Colombia: Magdalena, Nariño.
metathoracicus (de Saussure)
Polybia
metathoracica de Saussure, 1854, Ét. Fam. Vesp. 2: 198, female (in division
My) - [French Guiana:] “Cayenne”
(lectotype Paris); 1855, Ét. Fam. Vesp. 2.: pl. XXV fig. 1. -
Smith, 1857, Cat. Hym. Ins. Coll. Brit. Mus. 5, Vesp.: 130. - Dalla Torre, 1894, Cat. Hym. 9: 164 (cat.). -
Fox, 1898, Proc. Acad. Nat. Sci. Philad.: 449 (
Megacanthopus metathoracicus; Ducke,
1904, Bol. Mus. Goeldi, Pará 4: 360 (key), 362 (nest;
Mischocyttarus metathoracicus; Ducke, 1918, Rev. Mus.
Paul. 10: 353 [partim]. - ? Zikán,
1935, Arq. Inst. Biol. Veget. 1 (3): 155 (key), figs. 72, 74.
- Bequaert, 1933, Entomologica Am. 13: 118 (nest).- Richards, 1941, Proc. R.
Ent. Soc. London (B) 10: 125 (in subgenus Kappa,
ater group). - Bequaert, 1944, Bull.
Mus. Comp. Zool. 94: 262 (list). - Richards, 1945, Trans. R. Ent. Soc. London
95: 312 (key), 346, figs. 34, 38-40 (in subgenus Kappa, ater group). -
Zikán, 1949, Bol. Parq. Nac. Itatiaia 1: 39 (key), 203, fig. 138. - Richards,
1978, Soc. Wasps Am.: 292 (key), 307 (in subgenus Kappa; nest). - Overal, 1978, Bol. Mus. Para. Emílio Goeldi (N.
S.), Zool. 94: 10 (list). - Silveira, 2006, Zootaxa 1321: 15 (key), 71, figs.
66-68, 77, 79 (syns.: M. granadaenis Zikán, M. rectus Richards;
Mischocyttarus metathoracicus var. metathoracicus; Richards, 1945, Trans.
R. Ent. Soc. London 95: 313 (key), 346 (in subgenus Kappa, ater group)
Mischocyttarus
granadaënsis Zikán, 1949, Bol. Parq. Nac. Itatiaia
1: 39 (key), 202, figs. 137, 280, female - [
Mischocyttarus
rectus Richards, 1978, Soc. Wasps Am.: 292 (key),
306, female, ? male (in subgenus Kappa)
- “
Distr.: Colombia; Guyana; Suriname; French Guiana; Peru:
Loreto; Brazil: Amapá, Amazonas, Mato Grosso, Pará, Rio de Janeiro, Roraima;
Bolivia: Cochabamba, Santa Cruz.
mirus Silveira
Mischocyttarus
mirus Silveira, 2006, Zootaxa 1321: 3, 11 (key), 26, fig. 18, female (in
subgenus Kappa) - “Brazil, MG, Viçosa, Mata do Paraíso” (Curitiba).
Distr.: Brazil: Minas Gerais.
mocsaryi (Ducke)
Megacanthopus
Mocsáryi Ducke, 1909, Ann. Hist.-Nat. Mus. Natl. Hung. 7: 627, female, male
- [Brazil:] “Obidos (Bas Amazone)” (Paris).
Mischocyttarus mocsaryi; Ducke, 1918, Rev. Mus. Paul.
10: 351. - Richards, 1945, Trans. R. Ent. Soc. London 95: 341.
- Zikán, 1949, Bol. Parq. Nac. Itatiaia 1: 39 (key), 198. - Richards, 1978,
Soc. Wasps Am.: 290 (key), 300 (in subgenus Kappa).
- Nascimento and Overal, 1979, Bol. Mus. Para. Emílio Goeldi (N. S.) 95: 9
(list). - Silveira, 2006, Zootaxa 1321: 15 (key), 66, figs. 35-37 (in subgenus Kappa).
Distr.:
mysticus Silveira
Mischocyttarus
mysticus Silveira, 2006, Zootaxa 1321: 4, 17 (key),
102, figs. 82-83, female (in subgenus Kappa) - “
Distr.:
occiduus Silveira
Mischocyttarus
metathoracicus; Richards, 1978, Soc. Wasps Am.: 307
[nest; partim].
Mischocyttarus
occiduus Silveira, 2006, Zootaxa 1321: 4, 15 (key),
77, figs. 74, 80, female (in subgenus Kappa) - “
Distr.: Colombia: Cauca, Nariño.
paris Silveira
Mischocyttarus
paris Silveira, 2006, Zootaxa 1321: 3, 12 (key), 33, figs. 20-22, 24, male
(in subgenus Kappa) - “Peru, Madre de
Dios, Avipas, 400m” (New York).
Distr.: Peru:
Madre de Dios.
pertinax
Silveira
Mischocyttarus pertinax Silveira, 2006, Zootaxa 1321: 4, 18 (key), 103,
figs. 88-89, female, nest (in subgenus Kappa) - “Brazil, Pará, Bujarú” (Belem).
Distr.:
picturatus Bequaert
Polybia rufidens; Fox, 1898,
Proc. Acad. Nat. Sci. Philad.: 450 [partim].
Mischocyttarus
metathoracicus var. picturatus Bequaert, 1938, Proc. R. Ent. Soc. London 7: 135, female
- “
Mischocyttarus picturatus; Carpenter and
van der Vecht, 1991, Ann. Carnegie Mus. 60 (3): 216 (holotype is from Chapada;
paratype is M. bertonii; in subgenus Kappa). - Silveira, 2006, Zootaxa 1321:
16 (key), 84 (in subgenus Kappa).
Distr.: Brazil: Mato Grosso.
porantin Silveira
Mischocyttarus
porantin Silveira, 2006, Zootaxa 1321: 4, 16 (key), 85, figs. 91, 94, 96,
female, male, nest (in subgenus Kappa) - “Brazil, Pará, Bujarú” (holotype
female Belem).
Distr.:
pseudomimeticus (Schulz)
Polybia
pseudomimetica Schulz, 1904 (1903), Sber. Akad.
Bayer. Wiss. 33: 803, 810 (key), female - [
Megalacanthopus [!] pseudomimeticus;
Schulz, 1906, Spolia Hym. 234. - Ducke, 1907, Bol. Mus. Para. Emilo Goeldi 5: 190 (Megacanthopus; nest) [partim];
1910, Ann. Hist.-Nat. Mus. Natl. Hung. 8: 541.
Mischocyttarus pseudomimeticus;
Ducke, 1918, Rev. Mus. Paul.
10: 350 [partim]. - Richards, 1941,
Proc. R. Ent. Soc. London (B) 10: 125 (in subgenus Kappa, ater group); 1945, Trans. R. Ent. Soc. London 95: 312 (key), 348
(in subgenus Kappa, ater group); 1978, Soc. Wasps Am.: 290
(key), 300 (in subgenus Kappa). -
Overal, 1978, Bol. Mus. Para. Emílio Goeldi (N. S.), Zool. 94: 10
(list). - Sarmiento, 1994, Rev. Biol. Trop. 42 (1/2): 360 (Colombia: Meta;
list). - Silveira, 2006, Zootaxa 1321: 12 (key), 38, fig. 34 (in subgenus Kappa; nest; Bolivia; Brazil; Colombia; Peru).
Distr.: Colombia: Meta; Peru: Loreto; Brazil: Amazonas;
Bolivia: Pando.
santacruzi Raw
Mischocyttarus
santacruzi Raw, 2000, Rev. Bras. Ent. 17 (4): 941, female, male, nest (in
subgenus Kappa) - “1.6 km south-west
of the administrator’s house at Fazenda Baixão (15°09’S, 39°07’W) and 20 km
from the coast in the municipality of Ilheus, eastern Bahia state, Brazil”
(holotype female São Paulo). - Silveira, 2006, Zootaxa 1321: 14 (key), 63 (in
subgenus Kappa).
Distr.: Brazil: Bahia.
schunkei Zikán
Mischocyttarus “schunkei Zikán nov. spec.” Weyrauch, 1943, Bol. Mus. Hist. Nat.
Javier Prado 7: 69. Nomen nudum.
Mischocyttarus schunkei Zikán, 1949, Bol. Parq. Nac. Itatiaia 1: 38 (key), 196, female, male, nest - “
Distr.: Peru:
Junín.
socialis
(de Saussure)
Polybia ignobilis; de Saussure, 1854, Ét. Fam.
Vesp. 2.: pl. XXIV fig. 5, and footnote to explanation of plate.
Polybia atra; de Saussure, 1854, Ét. Fam. Vesp. 2.: explanation of pl. XXIV fig. 5, not 181 text (in
division Iota); 1854, Ét. Fam. Vesp.
2.: VV, footnote to revised explanation of pl. XXIV fig. 5. - von Ihering, 1904, Rev. Mus. Paul.
6: 190. Misidentification.
Polybia socialis de Saussure, 1854, Ét. Fam.
Vesp. 2: 177 text not pl. XXIV fig. 1, female (in division Iota) - “Le Brésil” (
Megacanthopus ater; Ducke, 1905, Bol.
Mus. Goeldi, Pará 4: 689 (syns.: Polybia
socialis de Saussure, Polistes
apicalis de Saussure, M. imitator
Ducke); 1906, Rev. Ent.,
Mischocyttarus ater; Ducke,
1913, Deutsch. Ent. Zeitschr.: 332 (syns.: P.
ignobilis Haliday, P. socialis de
Saussure, P. nigra de Saussure, M. imitator Ducke); 1918, Rev. Mus.
Paul. 10: 351 (Venezuela). - da Fonseca, 1926, Rev. Mus. Paul. 14: 175. -
Bequaert, 1933, Entomologica Am. 13: 118 (syn.: P. decepta Fox). - Zikán, 1935, Arq. Inst. Biol. Veget. 1 (3): 155 (key), figs. 78, 83, pl. 2 fig. 2, pl. 3 figs. 5-6
(nest). - Richards, 1940, Ann. Mag. Nat. Hist. (11) 5: 182 (compared to M. immarginatus n. sp.); 1941, Proc. R.
Ent. Soc. London (B) 10: 125 (in subgenus Kappa,
ater group); 1945, Trans. R. Ent.
Soc. London 95: 312 (key), 352, figs. 29, 36-37, pl. 3 figs. 161-165 (in
subgenus Kappa, ater group). - Zikán, 1949, Bol. Parq. Nac. Itatiaia 1: 40 (key),
205, figs. 405-406 [partim]. -
Hamilton, 1972, Annu. Rev. Ecol. Syst. 3: 225 (ethology). - Overal, 1978, Bol.
Mus. Para. Emílio Goeldi (N. S.), Zool. 94: 10 (list).
Mischocyttarus socialis; Bequaert, 1944
(1943), Ann. Ent. Soc. Am. 36: 721 [partim];
1944, Bull. Mus. Comp. Zool. 94: 264 (list); 1949 (1948), Bol. Ent.
Mischocyttarus
atramentarius Zikán, 1949, Bol. Parq. Nac. Itatiaia
1: 40 (key), 206, figs. 140-141, 349, 396, 398, 407, 414, female, male, nest -
[
Mischocyttarus atra; Overal, 1978, Bol.
Mus. Para. Emílio Goeldi (N. S.), Zool. 94: 10 (list).
Distr.: Trinidad (?); Brazil: Minas Gerais, Rio de
Janeiro, São Paulo, Santa Catarina, Paraná; Paraguay: Alto Paraná; Argentina:
Misiones.
tapuya (Schulz)
Polybia tapuya Schulz,
1905, Hym. Stud.: 133, male - [Brazil] “In silvis propre Belem do Pará”
(London).
Megacanthopus tapuya; Schulz, 1906, Spolia Hym.: 234.
- Ducke, 1907, Z. Hym. Dipt. 1907:
140; 1910, Ann. Hist.-Nat. Mus. Natl. Hung. 8: 541.
Mischocyttarus tapuya;
Ducke, 1918, Rev. Mus. Paul.
10: 353. - Richards, 1941, Proc. R. Ent. Soc. London (B) 10: 125 (in subgenus Kappa, ater group); 1945, Trans. R. Ent. Soc. London 95: 312 (key), 348,
fig. 34 (in subgenus Kappa, ater group). - Zikán, 1949, Bol. Parq. Nac. Itatiaia 1: 23 (key), 117. - Richards, 1978, Soc. Wasps
Am.: 289 (key), 293 (in subgenus Kappa).
- Silveira, 2006, Zootaxa 1321: 12 (key), 32, figs. 23, 25 (in subgenus Kappa).
Distr.:
timbira Silveira
Mischocyttarus
timbira Silveira, 2006, Zootaxa 1321: 4, 16 (key),
88, figs. 90, 93, 95, female, male, nest (in subgenus Kappa) - “Brazil,
Maranhão, Barreirinhas, Tabocas, 03º02’S 43º07’W” (holotype female Belem); also
from elsewhere in Maranhão.
Distr.:
tolensis Richards
Mischocyttarus
tolensis Richards, 1941, Proc. R. Ent. Soc. London
(B) 10: 125 (key), 128, female (in subgenus Kappa,
ater group) - “
Distr.: Costa Rica; Panama.
villarricanus Zikán
Mischocyttarus
villarricanus Zikán, 1935, Arq. Inst. Biol. Veget. 1 (3):
155 (key), 200, figs. 79-80, pl. 3 fig. 8, female, nest - “Villaricca (
Distr.:
zikaninus Richards
Mischocyttarus
pseudomimeticus; Ducke, 1918, Rev. Mus. Paul.
10: 350 [partim]. - Zikán, 1949, Bol.
Parq. Nac. Itatiaia 1: 37 (key), 191, figs. 128, 278, 319.
Mischocyttarus
zikaninus Richards, 1978, Soc. Wasps Am.: 289 (key), 295, female (in
subgenus Kappa) - “BRAZIL: Pará,
óbidos” (São Paulo). - Silveira, 2006, Zootaxa 1321: 3, 12 (key), 37, figs.
32-33 (in subgenus Kappa; male; Brazil).
Distr.: Brazil: Amazonas, Pará.
Subgenus
Megacanthopus Ducke
Megacanthopus Ducke, 1904, Bol. Mus. Goeldi 4: 320, 358, genus (9 species).
Type species: Megacanthopus collaris
Ducke, 1904, by subsequent designation of Bequaert, 1933, Entomologica Am. 13:
112.
Megalacanthopus Schulz, 1906, Spolia Hym. 1906: 234. Unjustified emendation.
belemensis Cooper
Mischocyttarus
belemensis Cooper, 1997, Ent. Mon. Mag. 133: 219
(key), 221, fig. 14, male, female (in subgenus Megacanthopus) - “
Distr.:
clavicornis Cooper
Mischocyttarus
clavicornis Cooper, 1997, Ent. Mon. Mag. 133: 219
(key), 220, figs. 2, 3, 6, 9, 11, male, female, nest (in subgenus Megacanthopus) - “
Distr.:
collarellus Richards
Mischocyttarus
collarellus Richards, 1940, Ann. Mag. Nat. Hist.
(11) 5: 177, female, male - “British Guiana: Mazaruni Settlement” (holotype
female
Distr.:
collaris (Ducke)
Megacanthopus
collaris Ducke, 1904, Bol. Mus. Goeldi, Pará 4: 359
(key), 361, female, male - [Brazil] “Oyapoc” (lectotype female São Paulo); also
from Belem and Óbidos. - Richards, 1978, Soc. Wasps Am.: 384 (designation of lectotype).
- Nascimento and Overal, 1979, Bol. Mus. Para. Emílio Goeldi (N. S.) 95:
9 (list). - Carpenter, 1999, Am. Mus. Novitat. 3259: 20 (notes
on types).
Mischocyttarus collaris; Ducke, 1918,
Rev. Mus. Paul. 10: 354. - da Fonseca, 1926, Rev. Mus. Paul. 14: 181 (compared
to ypiranguensis n. sp.). - Bequaert,
1933, Entomologica Am. 13: 118 (nest), 119 (ethology).- Zikán, 1935, Arq. Inst.
Biol. Veget. 1 (3): 153 (key), figs. 21, 24, 26. - Richards, 1940, Ann. Mag.
Nat. Hist. (11) 5: 177 (compared to M.
collarellus and M. superus n.
spp.); 1941, Proc. R. Ent. Soc. London (B) 10: 127 (in subgenus Megacanthopus, collaris group). - Bequaert, 1944, Bull. Mus. Comp. Zool. 94: 261
(list). - Richards, 1945, Trans. R. Ent. Soc. London 95: 319 (key), 416, figs.
78-80 (in subgenus Megacanthopus, collaris group). - Zikán, 1949, Bol.
Parq. Nac. Itatiaia 1: 18 (key), 76, figs. 37-38, 178, 304. - Richards, 1978,
Soc. Wasps Am.: 383 (key), 384 (in subgenus Megacanthopus).
- Overal, 1978, Bol. Mus. Para. Emílio Goeldi (N. S.), Zool. 94: 10
(list). - Sarmiento, 1994, Rev. Biol. Trop. 42 (1/2): 361 (list). - Cooper,
1997, Ent. Mon. Mag. 133: 219 (key), 222 (nest), figs. 4, 8, 12.
Mischocyttarus
collaris var. collaris; Richards, 1945, Trans. R. Ent. Soc.
London 95: 319 (key), 417 (in subgenus Megacanthopus,
collaris group).
Distr.: Colombia: Vaupés; Guyana; Ecuador: Napo; Brazil:
Amapá, Amazonas, Pará; Argentina: Misiones.
inexspectatus
Cooper
Mischocyttarus
inexspectatus Cooper, 1997, Ent. Mon. Mag. 133: 217 (key), 220, figs. 1, 5,
10, male, female, nest (in subgenus Megacanthopus)
- “COLOMBIA: Nariño, Barbacoas” (holotype male London).
Distr.: Colombia: Nariño.
malaris Richards
Mischocyttarus
malaris Richards, 1978, Soc. Wasps Am.: 383 (key), 388, female, male (in
subgenus Megacanthopus) - “PERU:
Loreto, Cerro Azul near Contamana, R. Ucayali, 200 m” (holotype female London);
also from French Guiana. - Cooper, 1997, Ent. Mon. Mag. 133: 217 (key), 222.
Distr.: French Guiana; Peru: Loreto.
melanopygus Richards
Mischocyttarus
collaris var. melanopygus
Richards, 1945, Trans. R. Ent. Soc. London 95: 319 (key), 419, female (in
subgenus Megacanthopus, collaris group) - “
Mischocyttarus melanopygus; Zikán, 1949,
Bol. Parq. Nac. Itatiaia 1: 17 (key), 72, figs. 32-33, 177, 358, 416-417. -
Richards, 1978, Soc. Wasps Am.: 383 (key), 387 (in subgenus Megacanthopus). - Sarmiento, 1994, Rev.
Biol. Trop. 42 (1/2): 361 (list). - Cooper, 1997, Ent. Mon. Mag. 133: 217
(key), 223; 1998, Ent. Mon. Mag. 134: 301 (mimicry).
Distr.: Colombia: Amazonas, Caquetá, Putumayo; Ecuador:
Morona-Santiago, Napo; Peru: Junín, Loreto; Bolivia: La Paz.
parallelogrammus Zikán
Polybia mexicana ?; Fox, 1898, Proc. Acad. Nat. Sci. Philad.: 451 [partim].
- Bequaert, 1933, Entomologica Am. 13: 142 (probably M. alfkenii).
Mischocyttarus
parallelogrammus Zikán, 1935, Arq. Inst. Biol.
Veget. 1 (3): 154 (key), 172, figs. 33, 35, 37, pl. 1 fig. 2, female, male,
nest - [
Mischocyttarus collaris var. parallelogrammus; Richards, 1940, Ann.
Mag. Nat. Hist. (11) 5: 177 (compared to dark var. of M. collarellus n. sp.); 1945, Trans. R. Ent. Soc. London 95: 319
(key), 418, pl. 3 fig. 180 (in subgenus Megacanthopus,
collaris group).
Mischocyttarus
parallelogrammoides Zikán, 1949, Bol. Parq. Nac.
Itatiaia 1: 236, sex and locality not stated (lectotype female Inst. Oswaldo
Cruz,
Distr.: Brazil: Minas Gerais, Rio de Janeiro, Santa
Catarina, São Paulo; Paraguay: Paraguarí.
saturatus Zikán
Mischocyttarus “collaris nov. var. saturatus Zikán” Weyrauch, 1943, Bol. Mus. Hist. Nat. Javier Prado
7: 68. Nomen nudum.
Mischocyttarus
collaris race saturatus Zikán,
1949, Bol. Parq. Nac. Itatiaia 1: 18 (key), 77, figs. 39-40, 287, 305, 343,
360, female, male - [Peru] “Tingo Maria, Rio Huallaga, 700 m” (lectotype male
Inst. Oswaldo Cruz,
Mischocyttarus saturatus; Richards,
1978, Soc. Wasps Am.: 383 (key), 387 (in subgenus Megacanthopus). - Sarmiento, 1994, Rev. Biol. Trop. 42 (1/2): 361 (
Distr.:
superus Richards
Mischocyttarus
superus Richards, 1940, Ann. Mag. Nat. Hist. (11)
5: 178, female, male - “British Guiana: Mazaruni Settlement” (holotype female
Distr.:
Guyana; Brazil: Pará.
Subgenus Mischocyttarus
de Saussure
acunai
Alayo
Mischocyttarus acuñai Alayo, 1972, Poeyana 94: 3, 4 (key), figs. 1B-E,
female, male - “Cuba ... Moa, Baracoa y Cupeyal (Yateras), ... de la provincia
de Oriente” (holotype male La Habana). - Richards, 1978, Soc. Wasps Am.: 310 (key), 327 (acunai;
in subgenus Monocyttarus). -
Silveira, 2002, Nat. Hist. Bull.
Distr.:
aripuanaensis Silveira
Mischocyttarus
aripuanaensis Silveira, 1998, Pap. Avuls. Zool.,
São Paulo 40 (23): 359, 360, figs. 1, 5, 7, 8a, 9, female (in subgenus Mischocyttarus) - “Brasil: MT, Aripuanã,
Reserva Humboldt” (
Distr.: Brazil: Mato Grosso.
cinerasceus Zikán
Mischocyttarus “cineraceus Zikán
nov. spec.” Weyrauch, 1943,
Bol. Mus. Hist. Nat. Javier Prado 7: 67.
Nomen nudum.
Mischocyttarus cinerasceus Zikán, 1949, Bol. Parq. Nac. Itatiaia 1:
13 (key), 49, fig. 158, female - [Peru] “Oxapampa, 1.300-1.500 m” (lectotype
Inst. Oswaldo Cruz, Rio de Janeiro); also from Oreja de Capelo and Chanchamayo.
- Richards, 1978, Soc. Wasps Am.: 275 (key), 279 (cinerascens [!]; designation of
lectotype; in subgenus Mischocyttarus).
- Colomo and Berta, 2005, Rev. Soc. Entomol. Argent. 64: 78 (note on types).
Distr.:
drewseni andinus Zikán
Mischocyttarus
“andinus Zikán
nov. spec.” Weyrauch, 1943,
Bol. Mus. Hist. Nat. Javier Prado 7: 67.
Nomen nudum.
Mischocyttarus drewseni race andinus
Zikán, 1949, Bol. Parq. Nac. Itatiaia 1: 14 (key),
52, fig. 164 - [
Mischocyttarus
litoralis race meridionalis
Zikán, 1949, Bol. Parq. Nac. Itatiaia 1: 14 (key), 54, figs. 15-16, 166,
female, male - “
Mischocyttarus
drewseni andinus; Richards, 1978, Soc. Wasps Am.:
275 (key), 277 (in subgenus Mischocyttarus).
- Sarmiento, 1994, Rev. Biol. Trop. 42 (1/2): 360 (list). - Colomo and Berta, 2005, Rev. Soc.
Entomol. Argent. 64: 78 (Myschocyttarus [!]; note on types).
Distr.:
drewseni
drewseni de Saussure
Mischocyttarus drewseni de Saussure, 1857, Ann. Soc. Ent. France (3) 5:
317, male - “Brèsil” (type depository unknown). - Dalla Torre, 1894, Cat. Hym.
9: 117 (drewsenii [!]; cat.); 1904,
Gen. Ins. 19: 85 (cat.). - Ducke, 1904, Bol. Mus. Goeldi, Pará 4: 318
(ecology), 364 (Brazil), fig. d. - Schrottky, 1904, Allg. Zeitschr. Ent. 9: 345
(Myschocyttarus [!]; Paraguay). -
Ducke, 1906, Rev. Ent., Caen 25: 10 (Brazil). - Bequaert, 1933, Entomologica
Am. 13: 118 (syns.: Megacanthopus
atriceps Cameron, Megacanthopus
rotundicollis Cameron). - Zikán, 1935, Arq. Inst.
Biol. Veget. 1 (3): 153 (key), figs. 1, 7, 12, pl. 2 fig. 7. - Richards, 1941,
Proc. R. Ent. Soc. London (B) 10: 124 (in subgenus Mischocyttarus, labiatus
group). - Bequaert, 1944, Bull. Mus. Comp. Zool. 94: 264 (list). - Richards,
1945, Trans. R. Ent. Soc. London 95: 310 (key), 328, figs. 6, 15 (in subgenus Mischocyttarus, labiatus group; syn.:? M.
mattogrossoensis Zikán). - Bequaert, 1949 (1948), Bol. Ent.
Megacanthopus
atriceps Cameron, 1912, Timehri (3) 2: 216 (key),
217, female - locality not stated [
Mischocyttarus
drewseni var.
drewseni; Richards, 1945, Trans. R. Ent. Soc. London 95: 310 (key).
Mischocyttarus
litoralis Zikán, 1949, Bol. Parq. Nac. Itatiaia 1:
14 (key), 53, figs. 14, 165, female, male - [
Mischocyttarus
montanus Zikán, 1949, Bol. Parq. Nac. Itatiaia 1:
14 (key), 54, 236 (“host” of montanoides
n. sp.), figs. 167, 397, female, male - [Brazil] “Itatiaia, desde 700 até 900
metros sôbre o nível do mar, Estado do Rio de Janeiro” (lectotype female Inst.
Oswaldo Cruz, Rio de Janeiro); also from Minas Gerais, São Paulo, Santa
Catarina, Rio Grande do Sul, Paraná; and Paraguay. - Richards, 1978, Soc. Wasps
Am.: 276 (designation of lectotype; syn. of M.
drewseni drewseni).
Mischocyttarus
montanus race nigrofemoratus
Zikán, 1949, Bol. Parq. Nac. Itatiaia 1: 15 (key), 56, male - [
Mischocyttarus
intermedius Zikán, 1949, Bol. Parq. Nac. Itatiaia
1: 15 (key), 57, figs. 17-18, 168, 355, female, male - “Itatiaia 400 a 900 m,
Estado no Rio de Janeiro, Brasil” lectotype female Inst. Oswaldo
Cruz,
Mischocyttarus
montanoides Zikán, 1949, Bol. Parq. Nac. Itatiaia
1: 236, sex and locality not stated (lectotype female Inst. Oswaldo Cruz,
Mischocyttarus
drewseni drewseni; Richards, 1978, Soc. Wasps Am.:
275 (key), 276 (syns.: M. litoralis Zikán,
montanus Zikán, montanus race nigrofemoratus Zikán,
intermedius Zikán, montanoides Zikán; in subgenus Mischocyttarus). - Almeida and Castro,
1991, Acta Biol. Par.,
Distr.: Colombia: Cauca; Venezuela:
Bolívar, D. F.; Guyana; Suriname; Peru: Cuzco, Loreto, San Martin; Brazil:
Amapá, Bahia, D. F., Espírito Santo, Goiás, Mato Grosso, Minas Gerais, Pará,
Paraná, Rio de Janeiro, Rio Grande do Sul, Santa Catarina, São Paulo; Bolivia:
Cochabamba, La Paz, Santa Cruz; Paraguay: Central, Cordillera, Paraguarí, San
Pedro, Concepción, Canindeyú, Alto Paraná; Argentina: Córdoba, Entre Rios,
Misiones, Tucumán (?).
drewseni gigas Richards
Mischocyttarus
drewseni var. gigas
Richards, 1945, Trans. R. Ent. Soc. London 95: 310 (key), 330, female, male
- “
Mischocyttarus
drewseni gigas; Willink, 1954 (1953), Acta Zool.
Lilloana 14: 321 (key), 322, figs. 5-7. - Richards, 1978, Soc. Wasps Am.: 275
(key), 278 (in subgenus Mischocyttarus).
Distr.:
gynandromorphus Richards
Mischocyttarus
gynandromorphus Richards, 1945, Trans. R. Ent. Soc.
London 95: 310 (key), 332, fig. 3, female, male (in subgenus Mischocyttarus, labiatus group) - “
Distr.: Peru: Junín; Brazil: Mato Grosso.
labiatus (Fabricius)
Zethus
labiatus Fabricius, 1804, Syst. Piez.: 284 -
“America meridionali” (lectotype female Kobenhavn).
Mischocyttarus labiatus;
de Saussure, 1853, Ét. Fam. Vesp. 2.: 20, pl. III
(female, male; French Guiana;
Distr.: Costa Rica; Panama; Colombia: Amazonas, Arauca,
Boyacá, Magdalena, Meta, Nariño, Putumayo, Valle, Vaupés; Venezuela: Bolívar,
D. F.; Trinidad; Guyana; Suriname; French Guiana; Ecuador: Napo; Peru:
Amazonas, Junín, Loreto, Madre de Dios; Brazil: Amazonas, D. F., Goiás, Mato
Grosso, Minas Gerais, Pará, Rio de Janeiro, Roraima; Bolivia: Beni, Pando,
Santa Cruz.
lindigi Richards
Mischocyttarus
lindigi Richards, 1945, Trans. R. Ent. Soc. London 95: 311 (key), 332,
female (in subgenus Mischocyttarus, labiatus group) - “COLOMBIA: Bogota”
(Berlin). - Zikán, 1949, Bol. Parq. Nac. Itatiaia 1: 216. -
Richards, 1978, Soc. Wasps Am.: 275 (key), 280 (in subgenus Mischocyttarus). - Sarmiento, 1994, Rev.
Biol. Trop. 42 (1/2): 360 (list). - Silveira, 1998, Pap. Avuls. Zool., São
Paulo 40 (23): 362, figs. 3-4 (redescription).
Distr.:
mattogrossoensis Zikán
Mischocyttarus
mattogrossoensis Zikán, 1935, Arq. Inst. Biol.
Veget. 1 (3): 153 (key), 156, figs. 4, 11, male - [
Mischocyttarus
drewseni var. steinbachi
Richards, 1945, Trans. R. Ent. Soc. London 95: 310 (key), 329, female, male
(in subgenus Mischocyttarus, labiatus group) - “
Distr.:
melanarius (Cameron)
Polybia melanaria Cameron,
1906, Invert. Pacif. 1: 153, female (in division Kappa) - “
Megacanthopus
longipetiolatus Cameron, 1912, Timehri (3) 2: 216
(key), 218, “female” [male] - “
Mischocyttarus melanarius; Richards,
1941, Proc. R. Ent. Soc. London (B) 10: 124 (in subgenus Mischocyttarus, labiatus
group); 1945, Trans. R. Ent. Soc. London 95: 311 (key), 322 (in subgenus Mischocyttarus, labiatus group; syn.: Megacanthopus
longipetiolatus Cameron). - Zikán, 1949, Bol. Parq. Nac. Itatiaia 1: 13
(key), 51, figs. 10-11, 160. - Richards, 1978, Soc. Wasps Am.: 275 (key), 280
(in subgenus Mischocyttarus; syn.: M. angustesulcatus Zikán). -
West-Eberhard et al., 1995, Hym. Costa
Rica: 585 (Costa Rica). - Rodríguez-Palafox, 1996, in Llorente-Bousquets
et al., Biodiv. Taxon. Biogeogr. Artrópod. México: 482 (list).
Mischocyttarus
angustesulcatus Zikán, 1949, Bol. Parq. Nac. Itatiaia 1: 12 (key), 44,
figs. 145-146, female, male - “Prietas, República de Honduras” (lectotype Inst.
Oswaldo Cruz, Rio de Janeiro). - Richards, 1978, Soc. Wasps Am.: 280
(designation of lectotype; syn. of M.
melanarius).
Distr.: Mexico: Chiapas*, Jalisco, Morelos, Tabasco,
Veracruz; Belize; Guatemala; Honduras; Costa Rica; Panama.
rotundicollis (Cameron)
Megacanthopus
rotundicollis Cameron, 1912, Timehri (3) 2: 216, “male” [female] - locality
not stated [Guyana] (London). - Bequaert, 1933, Entomologica Am. 13: 118 (syn.
of M. drewseni de Saussure).
Mischocyttarus
gracilis Zikán, 1935, Arq. Inst. Biol. Veget. 1 (3): 153
(key), 158, female, male, nest - [
Mischocyttarus
“fumigatus Zikán
nov. spec.” Weyrauch, 1943, Bol. Mus. Hist. Nat. Javier Prado 7: 67. Nomen
nudum.
Mischocyttarus rotundicollis; Richards,
1941, Proc. R. Ent. Soc. London (B) 10: 124 (in subgenus Mischocyttarus, labiatus
group; syn.: M. gracilis Zikán). -
Bequaert, 1944, Bull. Mus. Comp. Zool. 94: 263 (list). - Richards, 1945, Trans.
R. Ent. Soc. London 95: 311 (key), 330, figs. 7, 13-14, pl. 1 fig. 136, pl. 2
fig. 141 (in subgenus Mischocyttarus,
labiatus group). - Zikán, 1949, Bol.
Parq. Nac. Itatiaia 1: 13 (key), 47. - Willink, 1954 (1953), Acta Zool.
Lilloana 14: 321 (key), 324, figs. 1-4. - Vesey-Fitzgerald, 1956, Ent. Mon.
Mag. (4) 17: 165 (
Mischocyttarus
fumigatus Zikán, 1949, Bol. Parq. Nac. Itatiaia 1:
12 (key), 46, 236 (“host” of fumigatoides
n. sp.), figs. 5-6, 151-152, 323, female, male - [
Mischocyttarus
fumigatoides Zikán, 1949, Bol. Parq. Nac. Itatiaia
1: 236, sex and locality not stated (lectotype female Inst. Oswaldo Cruz,
Mischocyttarus
gracilioides Zikán, 1949, Bol. Parq. Nac. Itatiaia
1: 236, sex and locality not stated - (lectotype male Inst. Oswaldo Cruz,
Distr.: Colombia: Chocó, Meta, Valle;
Trinidad; Guyana; French Guiana; Ecuador: Napo, Pastaza, Zamora-Chinchipe;
Peru: Apurimac, Cuzco, Huánuco, Junín, Loreto, Pasco; Brazil: Amazonas, Bahia,
D. F., Espírito Santo, Rio de Janeiro, Goiás, Mato Grosso, Minas Gerais, Pará,
Paraná, Rio de Janeiro, Rondônia, Rio Grande do Sul, Santa Catarina, São Paulo;
Bolivia: La Paz, Santa Cruz; Paraguay: Cordillera, Paraguarí, Guairá, San
Pedro, Canindeyú; Uruguay; Argentina: Misiones.
smithii
de Saussure
Mischocyttarus smithii de Saussure, 1853, Ét. Fam. Vesp. 2: 21 - “Les
Amazones” (holotype male London). - Dalla Torre, 1894, Cat. Hym. 9: 117 (cat.);
1904, Gen. Ins. 19: 85 (Smithi [!];
cat.). - Zikán, 1935, Arq. Inst. Biol. Veget. 1 (3):
153 (smithi [!]; key), figs. 8, 13. -
Richards, 1941, Proc. R. Ent. Soc. London (B) 10: 124 (in subgenus Mischocyttarus, labiatus group). - Bequaert, 1944, Bull. Mus. Comp. Zool. 94: 263
(list). - Richards, 1945, Trans. R. Ent. Soc. London 95: 310 (key), 327, figs.
5, 18 (in subgenus Mischocyttarus, labiatus group; syn.: Megacanthopus violaceipennis Cameron). -
Zikán, 1949, Bol. Parq. Nac. Itatiaia 1: 15 (key), 58, figs. 19, 169, 300, 311
(Smithi [!]). - Richards, 1978, Soc.
Wasps Am.: 275 (key), 278 (in subgenus Mischocyttarus).
- Overal, 1978, Bol. Mus. Para. Emílio Goeldi (N. S.), Zool. 94: 10 (list). -
Almeida and Castro, 1991, Acta Biol. Par.,
Megacanthopus
violaceipennis Cameron, 1912, Timehri (3) 2: 216
(key), 217, “female” [male] - locality not stated [
Distr.:
tomentosus Zikán
Mischocyttarus
tomentosus Zikán, 1935, Arq. Inst. Biol. Veget. 1
(3): 153 (key), 155, figs. 3, 5, female - [
Mischocyttarus
“gibbicollis Zikán
nov. spec.” Weyrauch, 1943,
Bol. Mus. Hist. Nat. Javier Prado 7: 67.
Nomen nudum.
Mischocyttarus archidonaënsis Zikán, 1949, Bol. Parq. Nac. Itatiaia 1: 12 (key), 43, fig. 1, female - “Archidona, (
Mischocyttarus
gibbicollis Zikán, 1949, Bol. Parq. Nac. Itatiaia
1: 12 (key), 43, figs. 2, 144, female, male - [
Distr.:
Subgenus Monogynoecus
Richards
Monogynoecus Richards, 1941, Proc. Ent. Soc. Lond. (B)
10: 126, subgenus of Mischocyttarus
de Saussure.
Type species: Megacanthopus lecointei Ducke, 1904, by original designation.
carinulatus
Zikán
Mischocyttarus carinulatus Zikán, 1949, Bol. Parq. Nac. Itatiaia 1:
17 (key), 71, female - “Estado do Espírito Santo, Brazil” (São Paulo). -
Richards, 1978, Soc. Wasps Am.: 375 (key), 379 (in subgenus Monogynoecus). - Cooper, 1996, Ent. Mon.
Mag. 132: 24 (in subgenus Monogynoecus),
25 (key).
Distr.: Brazil: Bahia (?), Espírito Santo, Minas Gerais,
São Paulo*.
foveatus Richards
Mischocyttarus
foveatus Richards, 1941, Proc. R. Ent. Soc. London
(B) 10: 126 (key), 130, female, male (in subgenus Monogynoecus) - “BRITISH GUIANA: Essequibo R., 1st falls” (holotype
female
Distr.:
insolitus Zikán
Mischocyttarus
insolitus Zikán, 1949, Bol. Parq. Nac. Itatiaia 1:
16 (key), 59, figs. 20-21, 170-171, 312-313, 356, female, male - “Yaureté no rio Caiary, afluente do rio Uaupés, Estado do Amazonas, Brasil”
(lectotype male Inst. Oswaldo Cruz, Rio de Janeiro). - Richards, 1978, Soc.
Wasps Am.: 375 (key), 380 (in subgenus Monogynoecus).
- Carpenter and Wenzel, 1988, Psyche 95: 94. - Sarmiento, 1994, Rev. Biol.
Trop. 42 (1/2): 361 (
Distr.:
lecointei
guianensis Richards
Mischocyttarus lecointei var. guianensis
Richards, 1945, Trans. R. Ent. Soc. London 95: 319 (key), 409, figs. 74-75,
pl. 1 figs. 125-126, male, female, nest (in subgenus Monogynoecus) - “BRITISH GUIANA: north bank Essequibo R., Moraballi
Creek, Wallaba (Iperua) bush”
(holotype female London). - Zikán, 1949, Bol. Parq. Nac. Itatiaia 1: 218.
Mischocyttarus lecointei
guianensis; Richards, 1978, Soc.
Wasps Am.: 376 (key), 382 (in subgenus Monogynoecus).
- Cooper, 1996, Ent. Mon. Mag. 132: 24, 25 (key).
Distr.: Guyana.
lecointei lecointei (Ducke)
Polybia
surinamensis; Fox, 1898, Proc. Acad. Nat. Sci.
Philad.: 448 [partim].
Megacanthopus
lecointei Ducke, 1904, Bol. Mus. Goeldi, Pará 4:
359 (key), 361, female, male, nest - [Brazil] “Oyapoc” (lectotype male London);
also from Óbidos and Camahipy. - Richards, 1945, Trans. R. Ent. Soc. London 95:
408 (designation of lectotype); 1978, Soc. Wasps Am.: 381 (lectotype “here
designated”). - Nascimento and Overal, 1979, Bol. Mus. Para. Emílio
Goeldi (N. S.) 95: 9 (list). - Carpenter, 1999, Am. Mus. Novitat. 3259: 21 (notes on types).
Polybia ostenta du Buysson, 1908, Rev. Ent., Caen 27: 217, female
- “Guyane française: Nouveau Chantier du Bas Maroni” (Paris). - Ducke, 1910,
Ann. Hist.-Nat. Mus. Natl. Hung. 8: 541 (syn. of M. lecointei
Ducke). - Richards, 1945, Trans. R. Ent. Soc. London 95: 406 (? syn. of M. lecointei
Ducke).
Mischocyttarus lecointei; Ducke, 1918,
Rev. Mus. Paul. 10: 354. - da Fonseca, 1926, Rev. Mus. Paul.
14: 181 (compared to ypiranguensis n.
sp.). - Zikán, 1935, Arq. Inst. Biol. Veget. 1 (3): 153 (key), figs. 15, 20. -
Richards, 1940, Ann. Mag. Nat. Hist. (11) 5: 181 (compared to M. metoecus n. sp.); 1941, Proc. R. Ent.
Soc. London (B) 10: 126 (type of new subgenus Monogynoecus). - Bequaert, 1944, Bull. Mus. Comp. Zool. 94: 262
(list). - Richards, 1945, Trans. R. Ent. Soc. London 95: 319 (key), 406, pl. 3
figs. 156-160 (in subgenus Monogynoecus;
? syn.: Polybia ostenta du Buysson).
- Zikán, 1949, Bol. Parq. Nac. Itatiaia 1: 16 (key), 62, figs. 22, 172. -
Overal, 1978, Bol. Mus. Para. Emílio Goeldi (N. S.), Zool. 94: 10 (list). -
Carpenter and van der Vecht, 1991, Ann. Carnegie Mus. 60 (3): 215 (note on
material seen by Fox). - Sarmiento, 1994, Rev. Biol. Trop. 42 (1/2): 361 (Colombia; list).
Mischocyttarus lecointei
var. lecointei; Richards, 1945,
Trans. R. Ent. Soc. London 95: 319 (key), 407 (in subgenus Monogynoecus).
Mischocyttarus lecointei
lecointei; Richards, 1978, Soc. Wasps
Am.: 376 (key), 381 (in subgenus Monogynoecus).
- Cooper, 1996, Ent. Mon. Mag. 132: 23 (in subgenus Monogynoecus; nest), 24, 25 (key), 28 (Colombia; Ecuador; Peru),
fig. 5.
Distr.:
Colombia: Amazonas, Caquetá, Putumayo; Guyana; Suriname; French Guiana;
Ecuador: Napo; Peru: Loreto; Brazil: Amapá, Amazonas, Pará; Bolivia: Beni.
melanops Cooper
Mischocyttarus
melanops Cooper, 1996, Ent. Mon. Mag. 132: 23, 25
(key), 26, figs. 1-2, 4, 8, 10-14, male, female, nest - “
metoecus Richards
Mischocyttarus
metoecus Richards, 1940, Ann. Mag. Nat. Hist. (11)
5: 181, female, male - “British Guiana: Mazaruni Settlement” (holotype female
Distr.:
montei Zikán
Mischocyttarus
montei Zikán, 1949, Bol. Parq. Nac. Itatiaia 1: 16
(key), 65, figs. 173-174, 357, female, male - “Araçatuba, Estado de S. Paulo,
Brasil” (lectotype male Inst. Oswaldo Cruz,
Distr.: Brazil: São Paulo.
onorei Cooper
Mischocyttarus
onorei Cooper, 1996, Ent. Mon. Mag. 132: 23, 24
(key), 25, figs. 6-7, 9, 15, female, male, nest - “
Subgenus Omega
de Saussure
Omega de Saussure, 1854, Ét. Fam. Vesp. 2: 206, division of subgenus Polybia of genus Polybia
Lepeletier.
Type species: Polybia filiformis de Saussure, 1854, by monotypy.
Monacanthocnemis Ducke, 1905, Revue Ent. 24: 6, 8, 21, genus.
Type
species: Polybia filiformis de Saussure, 1854, by monotypy.
buyssoni
(Ducke)
Monacanthocnemis Buyssoni Ducke, 1906, Rev. Ent. 25: 9, female, nest
- [Brazil] “Rio de Janeiro” (lectotype London); 1910, Ann. Hist.-Nat. Mus. Natl. Hung. 8: 521. - Bequaert, 1933, Entomologica
Am. 13: 114 (typical Mischocyttarus).
- Richards, 1978, Soc. Wasps Am.: 373 (designation of lectotype). - Nascimento
and Overal, 1979, Bol. Mus. Para. Emílio Goeldi (N. S.) 95: 10 (list). -
Carpenter, 1999, Am. Mus. Novitat. 3259: 20 (notes on types).
Monacanthocnemis
buyssoni; Ducke, 1907, Bol. Mus.
Para. Emilo Goeldi 5: 183, pl. 3, fig. 8.
Mischocyttarus buyssoni; Ducke, 1918, Rev. Mus. Paul. 10: 355. - Zikán, 1935, Arq. Inst.
Biol. Veget. 1 (3): 155 (key), 201 (in subgenus Monacanthocnemis). - Richards, 1941, Proc. R. Ent. Soc. London (B)
10: 126 (in subgenus Omega); 1945,
Trans. R. Ent. Soc. London 95: 318 (key), 403, fig. 71 (in subgenus Omega). - Zikán, 1949, Bol. Parq. Nac. Itatiaia 1: 40 (key), 208. - Richards, 1978, Soc. Wasps
Am.: 369 (key), 373 (designation of lectotype; syns.: M. cristatus Zikán, M.
cristatoides Zikán; in subgenus Monacanthocnemis).
- Overal, 1978, Bol. Mus. Para. Emílio Goeldi (N. S.), Zool. 94: 10 (list). -
Almeida and Castro, 1991, Acta Biol. Par.,
Mischocyttarus
cristatus Zikán, 1935, Arq. Inst. Biol. Veget. 1
(3): 155 (key), 201, figs. 81-82, 84-85, pl. 2 fig. 5, female, male, nest (in
subgenus Monacanthocnemis) - [Brazil]
“Südhang des Itatiaya in 700 m Seehöhe” (lectotype female São Paulo). -
Richards, 1941, Proc. R. Ent. Soc. London (B) 10: 126 (in subgenus Omega); 1945, Trans. R. Ent. Soc. London
95: 318 (key), 401 (in subgenus Omega).
- W. Zikán and Wydodzinsky, 1948, Bol. Serv. Nac. Pesq. Agron. 4: 74 (list). -
Zikán, 1949, Bol. Parq. Nac. Itatiaia 1: 209, 236 (“host” of cristatoides n. sp.), figs. 142-143,
282-283, 310, 328-329, 411. - Richards, 1978, Soc. Wasps Am.: 373 (designation
of lectotype; syn. of M. buyssoni
(Ducke)). - Colomo and Berta, 2005, Rev. Soc. Entomol. Argent. 64: 79 (note on
types).
Mischocyttarus
cristatoides Zikán, 1949, Bol. Parq. Nac. Itatiaia
1: 236 -
Distr.:
chalucas Snelling
Mischocyttarus
chalucas Snelling, 1983, Pan-Pacif. Ent. 59: 277, figs.
8-11, female, male - “
Distr.:
filiformis (de Saussure)
Polybia filiformis de Saussure, 1854, Ét. Fam. Vesp. 2: 206, male
[female] (in division Omega) - “Le
Brésil” (holotype female London); 1855, Ét. Fam. Vesp. 2: pl. XXVI fig. 8. -
Dalla Torre, 1894, Cat. Hym. 9: 163 (cat.); 1904, Gen. Ins. 19: 77 (cat.). -
Zavattari, 1906, Boll. Mus. Zool. Anat. Comp. R. Univ. Torino 21 (529): 17.
Megacanthopus filiformis;
Ducke, 1904, Bol. Mus. Goeldi, Pará 4: 359 (key), 360 (nest; Brazil).
Monacanthocnemis filiformis;
Ducke, 1905, Bol. Mus. Goeldi, Pará 4: 687; 1906, Rev.
Ent.,
Mischocyttarus filiformes [!]; Ducke, 1918, Rev. Mus. Paul. 10: 355.
- Bequaert, 1933, Entomologica Am. 13: 114 (filiformis;
typical Mischocyttarus). - Zikán,
1935, Arq. Inst. Biol. Veget. 1 (3): 155 (key; in
subgenus Monacanthocnemis). -
Richards, 1941, Proc. R. Ent. Soc. London (B) 10: 126 (filiformis;
in subgenus Omega); 1945, Trans. R.
Ent. Soc. London 95: 318 (key), 402, fig. 70 (filiformis; in subgenus Omega). - Zikán, 1949, Bol. Parq. Nac.
Itatiaia 1: 207. - Richards, 1978, Soc. Wasps Am.: 369 (key), 370 (in subgenus Monacanthocnemis). - Overal, 1978, Bol.
Mus. Para. Emílio Goeldi (N. S.), Zool. 94: 10 (filiformis; list).
Distr.: Ecuador: Napo (?); Brazil: Pará; Bolivia: Beni
(?).
napoensis Richards
Mischocyttarus
napoensis Richards, 1978, Soc. Wasps Am.: 369
(key), 373, female (in subgenus Monacanthocnemis)
- “
Distr.:
omicron Richards
Mischocyttarus
omicron Richards, 1978, Soc. Wasps Am.: 369 (key),
372, female (in subgenus Monacanthocnemis)
- “BRAZIL: Mato Grosso, base camp, 12°50’S, 51°47’W” (London); also from
another locality in Mato Grosso; and Bolivia.
Distr.: Brazil: Mato Grosso; Bolivia: Beni.
punctatus (Ducke)
Megacanthopus
punctatus Ducke, 1904, Bol. Mus. Goeldi,
Pará 4: 360 (key), 363, female, nest - [Brazil] “Alcantara, Estado do Maranhão”
(Belém); 1907, Bol. Mus. Para. Emilo Goeldi 5: 189, pl. 3 fig.
9. - Overal, 1978, Bol. Mus. Para. Emílio Goeldi (N. S.), Zool. 94: 6
(designation of holotype as “lectotype”). - Nascimento and Overal, 1979, Bol.
Mus. Para. Emílio Goeldi (N. S.) 95: 10 (list). - Carpenter, 1999, Am.
Mus. Novitat. 3259: 22 (notes on type).
Mischocyttarus punctatus; Ducke, 1918, Rev. Mus. Paul.
10: 353, 370. - da Fonseca, 1926, Rev. Mus. Paul. 14: 183, pl. 1 fig. 3 (nest).
- Vesey-Fitzgerald, 1939, Proc. R. Ent. Soc. London (A) 14:
105 (mimicry; nest). - Richards, 1941, Proc. R. Ent. Soc. London (B) 10: 126
(in subgenus Omega); 1945, Trans. R.
Ent. Soc. London 95: 318 (key), 404, figs. 72-73 (in subgenus Omega). - Zikán, 1949, Bol. Parq. Nac.
Itatiaia 1: 118. - Richards, 1978, Soc. Wasps Am.: 369 (key), 370 (in subgenus Monacanthocnemis). - Overal, 1978, Bol.
Mus. Para. Emílio Goeldi (N. S.), Zool. 94: 10 (list). - Sarmiento,
1994, Rev. Biol. Trop. 42 (1/2): 361
(Colombia; list). - Rodríguez-Palafox, 1996, in Llorente-Bousquets et
al., Biodiv. Taxon. Biogeogr. Artrópod. México: 482 (list). -
Garcete-Barrett, 1999, Avispas Soc. Parag.: 24 (key, figs. 7a, 7b), 27
(Paraguay).
Distr.: Mexico (?); Colombia: Meta, Putumayo, Valle,
Vaupés; French Guiana*; Trinidad; Suriname; Brazil: Maranhão, Pará, Rio Grande
do Sul (?); Paraguay: Concepción.
vaqueroi Zikán
Mischocyttarus
vaqueroi Weyrauch, 1943, Bol. Mus. Hist. Nat. Javier Prado 7: 69. Nomen nudum.
Mischocyttarus vaqueroi Zikán, 1949, Bol. Parq. Nac. Itatiaia 1:
119, figs. 203, 291, female - “Pozuso, Peru” (Inst. Oswaldo Cruz, Rio de
Janeiro). - Richards, 1978, Soc. Wasps Am.: 369 (key), 371 (in subgenus Monacanthocnemis). - Sarmiento, 1994,
Rev. Biol. Trop. 42 (1/2): 361 (list). - West-Eberhard et al., 1995, Hym. Costa Rica: 585 (Costa Rica).
Distr.: Costa Rica; Colombia: Magdalena, Meta, Nariño,
Putumayo, Valle; Ecuador: Napo; Peru: Huánuco, Loreto; Brazil: Pará.
Subgenus Phi de
Saussure
Phi de Saussure, 1854, Ét. Fam. Vesp. 2: 183, division III of subgenus Polybia of genus Polybia
Lepeletier (10 species); at plates 23 and 24: division of genus Polybia.
Type species: Vespa phthisica
Fabricius, 1793, by subsequent designation of Bequaert, 1944, Bull. Mus. Comp.
Zool. 94: 292.
Monocyttarus Richards, 1978, Soc. Wasps Am.: vi; 18, 273, 307, subgenus of Mischocyttarus de Saussure.
Type species: Polybia flavitarsis de Saussure, 1854, by original designation.
alfkenii alfkenii (Ducke)
Megacanthopus
alfkenii Ducke, 1904, Bol. Mus. Goeldi, Pará 4: 360
(key), 362, female, male, nest - [Brazil] “nas mattas dos rios Villanova e
Camahipy (muncipio de Mazagão)” (lectotype female São Paulo); also from Oyapoc
and Óbidos; 1905, Bol. Mus. Goeldi, Pará 4: 688, pl. 4, figs. 24a-b;
1907, Bol. Mus. Para. Emilo Goeldi 5: 188. - Richards, 1978, Soc. Wasps Am.: 343 (designation of lectotype). - Nascimento and Overal,
1979, Bol. Mus. Para. Emílio Goeldi (N. S.) 95: 9 (list). - Carpenter,
1999, Am. Mus. Novitat. 3259: 19 (notes on types).
Mischocyttarus phthisicus var. alfkeni [!]; Ducke, 1913, Deutsch. Ent. Zeitschr.: 331.
Mischocyttarus alfkenii; Bequaert, 1933,
Entomologica Am. 13: 116 (this species was misidentified as M. indeterminabilis and M. bistriatus; compared to M.
cubensis). - Zikán, 1935, Arq. Inst. Biol. Veget. 1 (3): 153 (alfkeni [!]; key), figs. 22, 27; 1949,
Bol. Parq. Nac. Itatiaia 1: 160. - Richards, 1941, Proc. R. Ent. Soc. London
(B) 10: 126 (in subgenus Kappa, cubensis group); 1945, Trans. R. Ent.
Soc. London 95: 317 (key), 395, figs. 66, 68 (in subgenus Kappa, cubensis group)..
Mischocyttarus alfkenii var. alfkenii; Richards, 1945, Trans. R. Ent.
Soc. London 95: 318 (key), 395 (in subgenus Kappa,
cubensis group).
Mischocyttarus
japuraënsis Zikán, 1949, Bol. Parq. Nac. Itatiaia
1: 29 (key), 147, female - “Rio Japurá, Estado do Amazonas, Brasil” (
Mischocyttarus alfkenii alfkenii;
Richards, 1978, Soc. Wasps Am.: 313 (key), 344 (syn.: M. japuraensis Zikán; in subgenus Monocyttarus).
Distr.:
Brazil: Amapá, Amazonas, Pará.
alfkenii excrucians Richards
Mischocyttarus alfkenii; Bequaert, 1944,
Bull. Mus. Comp. Zool. 94: 264 (list).
Mischocyttarus
alfkenii var. excrucians
Richards, 1945, Trans. R. Ent. Soc. London 95: 318 (key), 396, pl. 1 fig.
131, female, male, nest (in subgenus Kappa,
cubensis group) - “COLOMBIA: Dept.
Meta, Restrepo, 500 m.” (holotype female
Mischocyttarus alfkenii excrucians;
Richards, 1978, Soc. Wasps Am.: 313 (key), 344 (in subgenus Monocyttarus). - Sarmiento, 1994, Rev.
Biol. Trop. 42 (1/2): 361 (list).
Distr.: Colombia: Meta, Putumayo; Venezuela: Aragua;
Guyana; Suriname; Brazil: Acre.
alfkenii trinitatis Richards
Mischocyttarus basimacula; Martorell,
1939, J. Agric. Univ. Puerto Rico 23: 230 (Venezuela: Aragua). Misidentification.
Mischocyttarus
alfkenii var. trinitatis Richards,
1945, Trans. R. Ent. Soc. London 95: 318 (key), 397, pl. 1 fig. 130, female,
male, nest (in subgenus Kappa, cubensis group) - “TRINIDAD: Quare R.
valley” (holotype female London); also from another locality in Trinidad; and
Venezuela; Guyana; Colombia. - Bequaert, 1949 (1948), Bol. Ent.
Mischocyttarus alfkenii trinitatis;
Richards, 1978, Soc. Wasps Am.: 313 (key), 344 (in subgenus Monocyttarus). - Sarmiento, 1994, Rev.
Biol. Trop. 42 (1/2): 361 (list).
Distr.: Colombia: Atlantico, Putumayo, Valle; Venezuela:
Aragua, Delta Amacuro, D. F.; Trinidad; Guyana.
alternatus
Zikán
Mischocyttarus alternatus Zikán, 1949, Bol. Parq. Nac.
Itatiaia 1: 130, male, female - “1.100 m, ... sul da Serra do Itatiaia, Estado
do Rio de Janeiro, Brasil” (lectotype female Inst. Oswaldo
Cruz,
Distr.:
angulatus angulatus Richards
Polybia indeterminabilis;
Cheesman, 1929, Trans. Ent. Soc. London 77: 145. Misidentification.
Mischocyttarus
angulatus Richards, 1945, Trans. R. Ent. Soc.
London 95: 317 (key), 386, figs. 63, pl. 1 fig. 132 (in subgenus Kappa, cubensis group). - Itô, 1984, J. Ethol. 2: 17-29 (ethology). -
Starr, 1988, Biotropica 20: 171 (
Mischocyttarus
angulatus var. angulatus; Richards, 1945, Trans. R. Ent. Soc.
London 95: 317 (key), 387, female, male (in subgenus Kappa, cubensis group) -
“
Mischocyttarus
angulatus var.
ictericus Richards, 1945, Trans. R. Ent. Soc. London 95: 317 (key), 389,
female, male (in subgenus Kappa, cubensis group) - “
Mischocyttarus
nevermanni Zikán, 1949, Bol. Parq. Nac. Itatiaia 1:
157, fig. 95, female - “San José da Costa Rica” (lectotype Inst. Oswaldo Cruz,
Mischocyttarus
flaviventris Zikán, 1949, Bol. Parq. Nac. Itatiaia 1: 164, figs. 245, 246,
female, male - “San José da Costa Rica” (lectotype male Inst. Oswaldo
Cruz,
Mischocyttarus
angulatus angulatus morph ictericus; Richards, 1978, Soc. Wasps Am.: 310 (key).
Mischocyttarus
angulatus angulatus morph angulatus; Richards, 1978, Soc. Wasps Am.: 310 (key).
Mischocyttarus angulatus angulatus;
Richards, 1978, Soc. Wasps Am.: 310 (key), 328 (syns.: M. nevermanni Zikán, M.
flaviventris Zikán; in subgenus Monocyttarus).
- Sarmiento, 1994, Rev. Biol. Trop. 42 (1/2): 361 (
Distr.: Mexico: Guerrero; El Salvador; Honduras; Costa
Rica; Panama; Colombia: Magdalena, Santander del Norte; adventive in
Martinique.
angulatus colombianus Richards
Mischocyttarus
angulatus var. colombianus
Richards, 1945, Trans. R. Ent. Soc. London 95: 317 (key), 388, female, male
(in subgenus Kappa, cubensis group) - “
Mischocyttarus angulatus colombianus;
West-Eberhard, 1975, Cespedesia 4 (16): 254 (
Distr.: Colombia: Antioquia, Boyacá, Cauca, Cundinamarca,
Huila, Magdalena, Meta, Valle; French Guiana (?); Bolivia (?).
aracatubaensis Zikán
Mischocyttarus
araçatubaënsis Zikán, 1949, Bol. Parq. Nac. Itatiaia 1: 32 (key), 162, fig.
412, female - “Araçatuba, Estado de São Paulo, Brasil” (Inst. Oswaldo
Cruz,
Distr.: Brazil: São Paulo.
araujoi Zikán
Mischocyttarus
araujoi Zikán, 1949, Bol. Parq. Nac. Itatiaia 1: 173, figs. 255-256, 391,
female, male - [Brazil] “Brumado, Muncípio de Amparo, Estado de S. Paulo”
(lectotype male Inst. Oswaldo Cruz, Rio de Janeiro). - Willink, 1954 (1953),
Acta Zool. Lilloana 14: 322 (key), 330, figs. 16-17 (Argentina: Entre Rios,
Misiones). - Richards, 1978, Soc. Wasps Am.: 314 (key), 355
(designation of lectotype; in subgenus Monocyttarus).
- Garcete-Barrett,
1999, Avispas Soc. Parag.: 8 (mimic of Agelaia
multipicta), 26 (key), 28 (
Distr.: Brazil: Rio de Janeiro, Santa Catarina, São
Paulo; Paraguay: Alto Paraná, Argentina: Entre Rios, Misiones.
bahiae Richards
Mischocyttarus
alfkenii var. bahiae
Richards, 1945, Trans. R. Ent. Soc. London 95: 318 (key), 398, female, male
(in subgenus Kappa, cubensis group) - “
Mischocyttarus bahiae; Zikán, 1949, Bol.
Parq. Nac. Itatiaia 1: 161, figs. 99-100, 242, 400. - Richards, 1978, Soc.
Wasps Am.: 313 (key), 345 (in subgenus Monocyttarus).
Distr.:
barbatulus Richards
Mischocyttarus
barbatulus Richards, 1978, Soc. Wasps Am.: 309 (key),
325, female, male (in subgenus Monocyttarus)
- “
Distr.:
barbatus Richards
Mischocyttarus
barbatus Richards, 1945, Trans. R. Ent. Soc.
Mischocyttarus
barbatus var. barbatus
Richards, 1945, Trans. R. Ent. Soc. London 95: 315 (key), 374, figs. 51, 59-60,
female, male (in subgenus Kappa, flavitarsis group) - “
Mischocyttarus barbatus var. cisandinus
Richards, 1945, Trans. R. Ent. Soc. London 95: 315
(key), 374, fig. 52, female (in subgenus Kappa,
flavitarsis group) - “
Distr.: Colombia: Cauca, Huila, Valle; Ecuador:
Morona-Santiago; Bolivia: Cochabamba.
basimacula basimacula (Cameron)
Polybia basimacula Cameron,
1906, Invert. Pacif. 1: 150, female - “Belize” (lectotype London).
Megacanthopus
indeterminabilis var. basimacula;
Ducke, 1910, Ann. Hist.-Nat. Mus. Natl. Hung. 8: 523, 539.
Mischocyttarus phthisicus var. basimacula; Ducke, 1913, Deutsch. Ent. Zeitschr.: 331.
Mischocyttarus basimacula; Bequaert, 1933, Entomologica
Am. 13: 117 (Guatemala; Honduras; Costa Rica; Panama; compared to M.
cubensis), 118 (nest). - Richards, 1941, Proc. R. Ent. Soc. London (B) 10: 126 (in subgenus Kappa,
cubensis group); 1945, Trans. R. Ent.
Soc. London 95: 317 (key), 391, 392 (designation of lectotype), figs. 65, 67,
pl. 1 figs. 127-129 (in subgenus Kappa,
cubensis group). - Zikán, 1949, Bol.
Parq. Nac. Itatiaia 1: 158, figs. 96-97, 239-240. - Itô, 1984, J. Ethol. 2:
17-29 (ethology). - Young, 1991, Brenesia 34: 51-61 (51, basimaculatus [!]; ecology). - Queller et al., 1992, J. Hym. Res. 1: 81-89 (genetic relatedness). -
West-Eberhard et al., 1995, Hym.
Mischocyttarus basimacula var. basimacula; Richards, 1945, Trans. R.
Ent. Soc. London 95: 318 (key), 391 (in subgenus Kappa, cubensis group).
Mischocyttarus
basimacula var. guatamalensis
Richards, 1945, Trans. R. Ent. Soc. London 95: 318 (key), 393, female, male
(in subgenus Kappa, cubensis group) - “
Mischocyttarus
basimacula from [form] guatamalensis; Richards, 1978, Soc. Wasps Am.: 313 (key).
Mischocyttarus basimacula basimacula; Richards, 1978, Soc. Wasps
Am.: 313 (key), 342 (in subgenus Monocyttarus).
- Sarmiento, 1994, Rev. Biol. Trop. 42 (1/2): 361 (list). - Rodríguez-Palafox, 1996, in
Llorente-Bousquets et al., Biodiv. Taxon. Biogeogr. Artrópod. México:
482 (list). - Hunt et al., 1999,
Biotropica 31 (1): 194 (migration).
Distr.: Mexico: San Luis Potosí, Tabasco, Veracruz,
Yucatán; Belize; Guatemala; El Salvador; Honduras; Nicaragua; Costa Rica;
Panama; Colombia: Nariño, Valle.
basimacula superpictus Richards
Mischocyttarus
basimacula var. superpictus
Richards, 1945, Trans. R. Ent. Soc. London 95: 318 (key), 393, female, male (in
subgenus Kappa, cubensis group) - “
Mischocyttarus basimacula superpictus; West-Eberhard, 1975,
Cespedesia 4 (16): 254 (superpectus
[!]). - Richards, 1978, Soc. Wasps Am.: 313 (key), 343 (in subgenus Monocyttarus). - Sarmiento, 1994, Rev.
Biol. Trop. 42 (1/2): 361 (list).
Distr.: Colombia: Boyacá, Cauca, Valle.
brackmanni Zikán
Mischocyttarus
brackmanni Zikán, 1949, Bol. Parq. Nac. Itatiaia 1: 129, 236 (“host” of brackmannoides n. sp.), figs. 76, 374,
male - [Brazil] “Muri, 1.100 m, Estado do Rio de Janeiro” (lectotype Inst. Oswaldo
Cruz, Rio de Janeiro). - Richards, 1978, Soc. Wasps Am.: 312 (key), 335
(designation of lectotype; syn.: M.
brackmannoides Zikán; in subgenus Monocyttarus).
Mischocyttarus
brackmannoides Zikán, 1949, Bol. Parq. Nac. Itatiaia 1:
236, sex and locality not stated (lectotype male Inst. Oswaldo Cruz,
Distr.: Brazil: Rio de Janeiro.
bruneri Bequaert and Salt
Mischocyttarus
bruneri Bequaert and Salt, 1931, Ann. Ent. Soc. Am. 24: 790, female -
“CUBA. Sierra Maestra, at 4,500 ft.” (
Distr.:
cabauna Zikán
Mischocyttarus
cabaúna Zikán, 1949, Bol. Parq. Nac.
Itatiaia 1: 139, fig. 224, female - “800 metros sôbre o nivel do mar, no Parque
Nacional do Itatiaia, Estado do Rio de Janeiro, Brasil” (lectotype Inst. Oswaldo Cruz, Rio de Janeiro). - Richards, 1978, Soc. Wasps Am.: 313
(key), 348 (designation of lectotype; in subgenus Monocyttarus).
Distr.: Brazil: Rio de Janeiro.
campestris Raw
Mischocyttarus
campestris Raw, 1985, Rev. Bras. Ent. 29: 107, 108, figs. 9, 11, 13, 15, 17
and 19, 110, female, male - [Brazil:] “Serra do Roncador...Brasília, D. F.” (holotype
female Rio de Janeiro). - Garcete-Barrett, 1999, Avispas Soc. Parag.: 28 (compared to M. marginatus (Fox)).
Distr.: Brazil: D. F.
cassununga (R. von Ihering)
Polybia cassununga R.
von Ihering, 1903, Ann. Soc. Ent. France 72: 146, female, male, nest - [Brazil]
“São João Climaco, à deux lieues de S. Paulo ... Sao Bernardo “ (lectotype
female São Paulo); 1904, Rev. Mus. Paul. 6: 207, pl. 3 figs. 12, 12a. - Dalla
Torre, 1904, Gen. Ins. 19: 77 (cat.). - Richards, 1978, Soc. Wasps
Am.: 363 (designation of lectotype). - Carpenter, 1999, Am. Mus. Novitat. 3259:
20 (notes on types).
Megacanthopus cassununga; Ducke, 1906,
Rev. Ent.,
Mischocyttarus cassununga;
Ducke, 1918, Rev. Mus. Paul. 10: 352. - Bequaert, 1933, Entomologica Am.
13: 118 (nest). - Zikán, 1935, Arq. Inst. Biol. Veget. 1 (3):
154 (key), 182, figs. 48-49, 54-55, pl. 1 fig. 5. - Richards, 1941, Proc. R.
Ent. Soc. London (B) 10: 126 (in subgenus Kappa,
flavitarsis group); 1945, Trans. R.
Ent. Soc. London 95: 315 (key), 376, figs. 54, 62 (in subgenus Kappa, flavitarsis group; syn.: M.
imitator Zikán). - Zikán, 1949, Bol. Parq. Nac. Itatiaia 1: 131, 236
(“host” of cassunungoides n. sp.),
figs. 77-80, 216-219, 307-308, 321, 332-333, 375, 379, 394, 399. - Richards,
1978, Soc. Wasps Am.: 315 (key), 363 (syns.: M. pirassununga Zikán, M.
curupira Zikán, M. cassunungoides Zikán;
in subgenus Monocyttarus). - Overal,
1978, Bol. Mus. Para. Emílio Goeldi (N. S.), Zool. 94: 10 (list). - Gobbi and
Simôes, 1988, An. Soc. Ent. Brasil 17 (2): 421-436 (ethology). - Almeida and
Castro, 1991, Acta Biol. Par.,
Mischocyttarus imitator Zikán, 1935, Arq. Inst. Biol.
Veget. 1 (3): 154 (key), 180, figs. 42, 44-46, pl. 1
fig. 1, female, male, nest - [Brazil] “Südabhang des Itatiaya in 700 m. ü. d.
M.” (lectotype female Inst. Oswaldo Cruz,
Mischocyttarus
pirassununga Zikán, 1949, Bol. Parq. Nac. Itatiaia
1: 133, figs. 81, 220, female, male - [Brazil] “Pirassununga, Estado de S.
Paulo” (lectotype male Inst. Oswaldo Cruz,
Mischocyttarus
curupira Zikán, 1949, Bol. Parq. Nac. Itatiaia 1:
133, 236 (“host” of curupiroides n.
sp.), fig. 376, male, female - [
Mischocyttarus
cassunungoides Zikán, 1949, Bol. Parq. Nac.
Itatiaia 1: 236, sex and locality not stated (lectotype male Inst. Oswaldo
Cruz,
Mischocyttarus
curupiroides Zikán, 1949, Bol. Parq. Nac. Itatiaia
1: 236, sex and locality not stated (lectotype male Inst. Oswaldo Cruz,
Distr.: Brazil: Bahia, Espírito Santo, Minas Gerais, Rio
de Janeiro, Santa Catarina, São Paulo; Paraguay: Canindeyú, Alto Paraná,
Itapúa; Argentina: Misiones.
catharinaensis Zikán
Mischocyttarus
catharinaënsis Zikán, 1949, Bol. Parq. Nac. Itatiaia 1: 33 (key), 175, fig.
348, female - “Nova Teutônia, sul de Santa Catarina, Brasil” (Inst. Oswaldo Cruz,
Distr.: Brazil: Santa Catarina.
cearensis Richards
Mischocyttarus
confirmatus var. cearensis Richards,
1945, Trans. R. Ent. Soc. London 95: 315 (key), 375, figs. 53, 61, female, male
(in subgenus Kappa, flavitarsis group) - “BRAZIL: Ceará,
Serra de Baturité, 600 m.” (holotype female
Mischocyttarus cearensis; Zikán, 1949,
Bol. Parq. Nac. Itatiaia 1:
138, figs. 85, 223, 309. - Richards, 1978, Soc. Wasps
Am.: 315 (key), 362 (in subgenus Monocyttarus).
Distr.:
chapadae (Fox)
Polybia
chapadae Fox, 1898, Proc. Acad. Nat. Sci. Philad.:
453, female - “
Mischocyttarus chapadae;
Richards, 1941, Proc. R. Ent. Soc. London (B) 10: 126 (in subgenus Kappa, flavitarsis group); 1945, Trans. R. Ent.
Soc. London 95: 315 (key), 369 (in subgenus Kappa,
flavitarsis group). - Zikán, 1949,
Bol. Parq. Nac. Itatiaia 1: 214. - Richards, 1978, Soc. Wasps Am.: 310 (key),
331 (in subgenus Monocyttarus).
Distr.:
commixtus Richards
Mischocyttarus
“spadiceus Zikán
nov. spec.” Weyrauch, 1943, Bol. Mus. Hist. Nat. Javier Prado 7: 68. Nomen
nudum.
Mischocyttarus
commixtus Richards, 1945, Trans. R. Ent. Soc.
London 95: 313 (key), 371, female (in subgenus Kappa, flavitarsis group)
- “
Mischocyttarus
spadiceus Zikán, 1949, Bol. Parq. Nac. Itatiaia
1: 153, fig. 236, female - “Oreja de Capello, 1.600 m, (Peru)” (Inst. Oswaldo Cruz,
Distr.:
confirmatus Zikán
Mischocyttarus
confirmatus Zikán, 1935, Arq. Inst. Biol. Veget. 1
(3): 154 (key), 164, male, nest - [
Mischocyttarus
confirmatus var. confirmatus; Richards,
1945, Trans. R. Ent. Soc. London 95: 315 (key), 376 (in subgenus Kappa, flavitarsis group).
Mischocyttarus
confirmatoides Zikán, 1949, Bol. Parq. Nac.
Itatiaia 1: 236, sex and locality not stated (lectotype male Inst. Oswaldo
Cruz,
Distr.: Brazil: Rio de Janeiro.
consimilis
Zikán
Mischocyttarus paraguayensis; Richards, 1941, Proc. R. Ent. Soc. London (B) 10: 126 (in subgenus Kappa,
flavitarsis group); 1945, Trans. R.
Ent. Soc. London 95: 315 (key), 379, fig. 55 (in subgenus Kappa, flavitarsis
group). Misidentification.
Mischocyttarus
consimilis Zikán, 1949, Bol. Parq. Nac. Itatiaia 1:
29 (key), 148, 236 (“host” of consimiloides
n. sp.), figs. 90-91, 230-231, 393, female, male - “Mbovevo, Paraguai”
(lectotype male
? Mischocyttarus
schrottkyi Zikán, 1949, Bol. Parq. Nac. Itatiaia 1: 159, figs. 98,
241, female - “Puerto Bertoni, (Paraguai)” (São Paulo).
Mischocyttarus
consimiloides Zikán, 1949, Bol. Parq. Nac. Itatiaia 1:
236, sex and locality not stated (lectotype female Inst. Oswaldo Cruz,
? Mischocyttarus
alfkeni [!] bahiae; Willink, 1954
(1953), Acta Zool. Lilloana 14: 322 (key), 336 (
Distr.:
costalimai Zikán
Mischocyttarus
costa-limai Zikán, 1949, Bol. Parq. Nac. Itatiaia 1: 180, figs. 117-118,
265-266, 401, female, male - “Itatiaia, 700 m. Estado doRio de Janeiro, Brasil”
(lectotype male Inst. Oswaldo Cruz,
Distr.:
Brazil: Rio de Janeiro.
costaricensis Richards
Mischocyttarus
costaricensis Richards, 1945, Trans. R. Ent. Soc.
London 95: 316 (key), 390, fig. 64, female, male (in subgenus Kappa, cubensis group) - “
Distr.: Guatemala; Costa Rica; Panama.
crypticus Zikán
Mischocyttarus
crypticus Zikán, 1949, Bol. Parq. Nac. Itatiaia 1: 141, 236 (“host” of crypticoides n. sp.), figs. 86, 226-227,
370, 385, female, male - “Itatiaia, 700 m, Estado do Rio de Janeiro, Brasil”
(lectotype male Inst. Oswaldo Cruz, Rio de Janeiro). - Richards, 1978, Soc.
Wasps Am.: 315 (key), 366 (designation of lectotype; syn.: M. crypticoides Zikán; in subgenus Monocyttarus).
Mischocyttarus
crypticoides Zikán, 1949, Bol. Parq. Nac. Itatiaia 1: 236,
sex and locality not stated (lectotype male Inst. Oswaldo Cruz,
Distr.: Brazil: Rio de Janeiro.
cryptobius Zikán
Mischocyttarus
cryptobius Zikán, 1935, Arq. Inst. Biol. Veget. 1 (3): 155 (key), 186,
female, male, nest - [Brazil] “Südabhange des Itatiaya (bei Kilometer 6,25, in
einer Höhe von etwa 1200 Metern)” (lectotype male Inst. Oswaldo Cruz, Rio de
Janeiro). - W. Zikán and Wydodzinsky, 1948, Bol. Serv. Nac. Pesq. Agron. 4: 74
(list). - Zikán, 1949, Bol. Parq. Nac. Itatiaia 1: 143, figs. 87, 386. -
Richards, 1978, Soc. Wasps Am.: 315 (key), 366 (in subgenus Monocyttarus).
Distr.:
cubensis
(de Saussure)
Polybia cubensis de Saussure, 1854, Ét. Fam. Vesp. 2: 202, pl XXV
figs. 5-6, female (in division My) -
“Les Antilles, Cuba” (lectotype Paris); 1856, in La Sagra, Hist. Phys.
Pol. Nat. Cuba, Segunda Parte 7: 324; 1857, in La Sagra, Hist. Phys.
Pol. Nat. Cuba, Segunda Parte 7: 774. - Smith, 1857, Cat. Hym. Ins. Coll. Brit. Mus. 5, Vesp.: 130 (cat.). - Cresson, 1865, Proc.
Entomol. Soc. Philadelphia 4: 167. - Gundlach, 1886, Contrib. Entomol. Cubana
2: 156. - Fox, 1890, Entomol. News 1: 93. - Dalla Torre, 1894, Cat. Hym. 9: 163
(cat.). - Ashmead, 1894, Psyche 7: 76; 1896, Bull. Labor. Nat. Sci.
Polybia phthisica; Cresson, 1865, Proc. Entomol. Soc. Philadelphia 4: 167. - Gundlach, 1886, Contrib. Entomol. Cubana
2: 155. Misidentification.
Megacanthopus
cubensis; Ducke, 1908, Dts. Entomol. Zeitschr.: 698
(male). - Meade-Waldo, 1911, Ann. Mag. Nat. Hist. (8) 7: 111.
Megacanthopus indeterminabilis var. cubensis;
Ducke, 1910, Ann. Hist.-Nat. Mus. Natl. Hung. 8: 540.
Mischocyttarus cubensis; Richards, 1978,
Soc. Wasps Am.: 308 (key), 317 (in subgenus Monocyttarus).
- Carpenter, 1999, Am. Mus. Novitat. 3259: 21 (note on types).
Distr.:
declaratus Zikán
Mischocyttarus
declaratus Zikán, 1935, Arq. Inst. Biol. Veget. 1
(3): 155 (key), 162, figs. 17-19, pl. 3 fig. 9, female, male, nest - [
Distr.:
Brazil: Rio de Janeiro.
duidensis
Richards
Mischocyttarus duidensis Richards, 1945, Trans. R. Ent. Soc. London 95: 314
(key), 370, figs. 48-49, female (in subgenus Kappa, flavitarsis group)
- “VENEZUELA: Mt. Duida” (Cambridge). - Bequaert, 1949 (1948), Bol. Ent.
Distr.:
ecuadorensis Zikán
Mischocyttarus
ecuadorensis Zikán, 1949, Bol. Parq. Nac. Itatiaia
1: 31 (key), 156, female - “Equador” (
Distr.:
extinctus Zikán
Mischocyttarus
extinctus Zikán, 1935, Arq. Inst. Biol. Veget. 1 (3):
155 (key), 183, figs. 50 (exinctus
[!]), 52, pl. 2 fig. 3, female, nest - [
Mischocyttarus
paranaënsis Zikán, 1949, Bol. Parq. Nac. Itatiaia
1: 140, fig. 225, female - [
Mischocyttarus
scotophilus Zikán, 1949, Bol. Parq. Nac. Itatiaia
1: 144, 236 (“host” of scotophiloides
n. sp.), figs. 88, 228, 387, female, male - “Itatiaia, 700 m, Estado do
Mischocyttarus
scotophiloides Zikán, 1949, Bol. Parq. Nac.
Itatiaia 1: 236, sex and locality not stated (lectotype male Inst. Oswaldo
Cruz,
Distr.:
fisheri Snelling
Mischocyttarus
fisheri Snelling, 1970, Contrib. Sci. 137:
1, 16, figs. 4d-f, female (in subgenus Kappa)
- [Mexico] “La Laguna, 5500’, Terr. Sur Baja California” (Los Angeles). -
Richards, 1978, Soc. Wasps Am.: 309 (key), 321 (in subgenus Monocyttarus). - Rodríguez-Palafox,
1996, in Llorente-Bousquets et al., Biodiv. Taxon. Biogeogr.
Artrópod. México: 482 (list).
Distr.: Mexico: Baja California Sur.
flavicornis flavicornis Zikán
Mischocyttarus
flavicornis Zikán, 1935, Arq. Inst. Biol. Veget. 1 (3):
154 (key), 168, female - [
Mischocyttarus basimacula var. flavicornis; Richards, 1945, Trans. R.
Ent. Soc. London 95: 317 (key), 394 (in subgenus Kappa, cubensis group).
Mischocyttarus
flavicornis flavicornis; Richards, 1978, Soc. Wasps
Am.: 313 (key), 341 (in subgenus Monocyttarus).
- Almeida and Castro, 1991, Acta Biol. Par.,
Distr.: Brazil: Goiás, Maranhão, Mato Grosso, Pará.
flavicornis nigricornis Zikán
Mischocyttarus “nigricornis Zikán nov. spec.” Weyrauch,
1943, Bol. Mus. Hist. Nat. Javier Prado 7: 68. Nomen
nudum.
Mischocyttarus
flavicornis race nigricornis
Zikán, 1949, Bol. Parq. Nac. Itatiaia 1: 154, female, male - “Santa Ana, Valle
de Rubamba, 1.100 metros, (Peru)” [Urubamba] (lectotype male Inst. Oswaldo Cruz,
Mischocyttarus flavicornis variety rufescens Zikán, 1949, Bol. Parq.
Nac. Itatiaia 1: 155, female, male - “Satipo, (Peru)” (lectotype male Inst. Oswaldo Cruz,
Mischocyttarus basimacula var. flavicornis; Richards, 1945, Trans. R.
Ent. Soc. London 95: 394 [partim].
Mischocyttarus
flavicornis nigricornis; Richards, 1978, Soc. Wasps
Am.: 313 (key), 341 (syn.: M. flavicornis var. rufescens Zikán; in
subgenus Monocyttarus). - Raw, 1989,
Maracá Rainfor. Proj., Invert. Limnol., Prelim. Rep.: 31 (compared to M. maracaensis n. sp.). - Sarmiento,
1994, Rev. Biol. Trop. 42 (1/2): 361 (list).
Distr.: Colombia: Arauca, Meta, Nariño; Ecuador: Manabi,
Napo; Peru: Cuzco, Huánuco, Junín, Loreto, Pasco; Bolivia: Beni.
flavitarsis
(de Saussure)
Polybia flavitarsis de Saussure, 1854, Ét. Fam.
Vesp. 2: 199, female (in division My)
- “La Californie” (
Megacanthopus flavitarsis; Ducke, 1907,
Bol. Mus. Para. Emilo Goeldi 5: 190; 1908, Deutsch. Ent. Z.: 698 (
Mischocyttarus flavitarsis; Essig, 1926,
Ins. West. North Am.: 888, fig. 750 (in subgenus Myrapetra). -
Mischocyttarus flavitarsis var. flavitarsis; Bequaert, 1933, Ent. Am.
(N. S.) 13: 123 (key), 125, 126 (nest), 127, pl. XXIX figs. 3-4 (
Mischocyttarus flavitarsis var. centralis
Bequaert, 1933, Ent. Am. (N. S.) 13: 123 (key),
125, 126-127 (stylopization), 129, female, male - [
Mischocyttarus
flavitarsis var. idahoensis Bequaert, 1933, Ent. Am. (N. S.) 13: 123 (key), 125,
133, female, male - [
Mischocyttarus flavitarsis var. kaibabensis
Bequaert, 1933, Ent. Am. (N. S.) 13: 122 (key),
125, 133, female - [
Mischocyttarus
flavitarsoides Zikán, 1949, Bol. Parq. Nac.
Itatiaia 1: 236, sex and locality not stated (Inst. Oswaldo Cruz,
Mischocyttarus flavitarsis flavitarsis; Bohart, 1951, in Muesebeck, et al., Hym. Am. N. of
Mischocyttarus flavitarsis centralis; Bohart, 1951, in Muesebeck, et al., Hym. Am. N. of
Mischocyttarus flavitarsis idahoensis; Bohart, 1951, in Muesebeck, et al., Hym. Am. N. of
Mischocyttarus flavitarsis kaibabensis; Bohart, 1951, in Muesebeck, et al., Hym. Am. N. of
Distr.:
flavoniger Zikán
Mischocyttarus
“flavoniger Zikán
nov. spec.” Weyrauch, 1943,
Bol. Mus. Hist. Nat. Javier Prado 7: 68.
Nomen nudum.
Mischocyttarus flavoniger Zikán, 1949, Bol. Parq. Nac. Itatiaia 1:
170, female - “Peru, (Vale de Chanchamayo, 800 m...)” (lectotype Inst. Oswaldo
Cruz, Rio de Janeiro); also from Oreja de Capelo. - Richards, 1978, Soc. Wasps Am.: 314 (key), 358 (designation of lectotype; in subgenus Monocyttarus). - Cooper, 1998, Ent. Mon.
Mag. 134: 301 (mimicry). - Colomo and Berta, 2005, Rev. Soc. Entomol. Argent. 64: 79 (note on types).
Distr.: Peru:
Junín; Bolivia: La Paz (?).
fluminensis
Zikán
Mischocyttarus fluminensis Zikán, 1949, Bol. Parq. Nac.
Itatiaia 1: 169, figs. 251, 388, female - [Brazil] “Rio de Janeiro, D. F.”
(Inst. Oswaldo Cruz,
Distr.: Brazil: Rio de Janeiro.
gilvus Zikán
Mischocyttarus
gilvus Zikán, 1949, Bol. Parq. Nac. Itatiaia 1: 31 (key), 159, male -
“Puerto Bortoni, (Paraguai)” (São Paulo). - Richards, 1978, Soc. Wasps Am.: 313 (key), 346 (in subgenus Monocyttarus). - Garcete-Barrett, 1999, Avispas Soc. Parag.: 26 (key), 28.
Distr.: Brazil*: São Paulo; Paraguay: Alto Paraná;
Argentina: Misiones (?).
haywardi Willink
Mischocyttarus
haywardi Willink, 1954 (1953), Acta Zool. Lilloana
14: 317, 322 (key), 327, figs. 8-9, female (in subgenus Kappa) - “Pto. 17 de Octubre
(ex Bemberg),
Distr.:
hirsutus Richards
Mischocyttarus
hirsutus Richards, 1945, Trans. R. Ent. Soc. London
95: 315 (key), 363 (in subgenus Kappa,
flavitarsis group); 1978, Soc. Wasps
Am.: 310 (key), 331 (in subgenus Monocyttarus).
- Sarmiento, 1993, Bol. Mus. Ent. Univ. Valle 1 (2): 6 (nest;
Mischocyttarus
hirsutus var. hirsutus
Richards, 1945, Trans. R. Ent. Soc. London 95: 315 (key), 364, female, male (in
subgenus Kappa, flavitarsis group) - “
Distr.:
hirtulus Zikán
Mischocyttarus
“furvus Zikán
nov. spec.” Weyrauch, 1943,
Bol. Mus. Hist. Nat. Javier Prado 7: 68.
Nomen nudum.
Mischocyttarus hirtulus Zikán, 1949, Bol. Parq. Nac. Itatiaia 1: 27 (key), 136, 236 (“host” of hirtuloides n. sp.), figs. 83-84, female, male - [Peru]
“Huacapistana, 1.800 m” (holotype male Inst. Oswaldo Cruz, Rio de Janeiro);
also from Carpapata. - Richards, 1978, Soc. Wasps Am.: 309 (key), 323 (syn.: M. hirtuloides Zikán; in subgenus Monocyttarus).
Mischocyttarus
hirtuloides Zikán, 1949, Bol. Parq. Nac. Itatiaia
1: 236, sex and locality not stated (lectotype female Inst. Oswaldo Cruz,
Distr.:
imeldai Zikán
Mischocyttarus
imeldai Weyrauch, 1943, Bol. Mus. Hist. Nat. Javier Prado 7: 69. Nomen
nudum.
Mischocyttarus imeldai Zikán, 1949, Bol. Parq. Nac. Itatiaia 1:
152, figs. 235, 295, female - “Peru (Vale Chanchamayo, 1.400 m)” (Inst. Oswaldo Cruz, Rio de Janeiro). - Richards, 1978, Soc. Wasps Am.: 312
(key), 333 (in subgenus Monocyttarus).
Distr.: Peru: Junín; Bolivia: La Paz.
inca Zikán
Mischocyttarus silvaticus
race inca Zikán, 1949, Bol. Parq.
Nac. Itatiaia 1: 191, female - “Peru” (São Paulo).
Mischocyttarus inca; Richards, 1978,
Soc. Wasps Am.: 310 (key), 326 (in subgenus Monocyttarus).
Distr.: Peru.
infrastrigatus Zikán
Mischocyttarus
infrastrigatus Zikán, 1949, Bol. Parq. Nac.
Itatiaia 1: 175, 236 (“host” of infrastrigatoides
n. sp.), figs. 112, 257, 390, female, male - [Brazil] “Itatiaia, 700 e 900 m,
Estado do Rio de Janeiro” (lectotype male Inst. Oswaldo Cruz, Rio de Janeiro).
- Richards, 1978, Soc. Wasps Am.: 314 (key), 360 (designation of lectotype;
syns.: M. ornatulus Zikán, M. infrastrigatoides Zikán; in subgenus Monocyttarus).
Mischocyttarus
ornatulus Zikán, 1949, Bol. Parq. Nac. Itatiaia
1: 177, figs. 114, 260-261, 392, female, male - [Brazil] “Itatiaia, 700 m,
Estado do Rio de Janeiro” (lectotype male Inst. Oswaldo Cruz,
Mischocyttarus
infrastrigatoides Zikán, 1949, Bol. Parq. Nac.
Itatiaia 1: 236, sex and locality not stated (lectotype female Inst. Oswaldo
Cruz,
Distr.: Brazil: Rio de Janeiro, São Paulo.
itatiayaensis Zikán
Mischocyttarus
itatiayaensis Zikán, 1935, Arq. Inst. Biol. Veget. 1 (3): 154 (key), 190,
figs. 58, 61-62, pl. 3 fig. 1, female, nest - [Brazil] “etwa 900 m Seehöhe am
Südabhang des Itatiaya” (lectotype Inst. Oswaldo Cruz, Rio de Janeiro); 1949, Bol.
Parq. Nac. Itatiaia 1: 179, figs. 115-116, 263-264, 389. - Richards, 1978, Soc.
Wasps Am.: 314 (key), 353 (designation of lectotype; in subgenus Monocyttarus).
Distr.: Brazil: Rio de Janeiro.
lanei Zikán
Mischocyttarus
lanei Zikán, 1949, Bol. Parq. Nac. Itatiaia 1: 25 (key), 127, figs. 74,
213, female - [Brazil] “Campos da Serra, Estado de S. Paulo” (São Paulo). -
Richards, 1978, Soc. Wasps Am.: 313 (key), 347 (in subgenus Monocyttarus).
Distr.: Brazil: São Paulo.
lilae Willink
Mischocyttarus
lilae Willink, 1954 (1953), Acta Zool. Lilloana 14:
317, 322 (key), 331, figs. 10-12, female, male (in subgenus Kappa) - [Argentina] “Amaicha del Valle
(2000 m s. n. m.), Tucumán” (holotype female Tucumán); also from Córdoba;
Salta; and Bolivia: Santa Cruz. - Richards, 1978, Soc. Wasps Am.: 314 (key),
352 (in subgenus Monocyttarus). - Colomo
and Berta, 2005, Rev. Soc. Entomol. Argent. 64: 80 (note on types).
Distr.: Bolivia: Santa Cruz; Argentina: Catamarca,
Córdoba, Corrientes, Salta, Tucumán.
lules Willink
Mischocyttarus
lules Willink, 1954 (1953), Acta Zool. Lilloana 14:
317, 322 (key), 334, figs. 13-15, female (in subgenus Kappa) - “Quebrada de Lules, Dpto. Famaillá, Tucumán, Argentina”
(Tucumán). - Richards, 1978, Soc. Wasps Am.: 314 (key), 352 (in subgenus Monocyttarus). - Colomo and Berta, 2005,
Rev. Soc. Entomol. Argent. 64: 81 (note on types).
Distr.: Argentina: Tucumán.
mamirauae Raw
Mischocyttarus
mamirauae Raw, 1998 (1996), Bol. Mus. Para. Emílio
Goeldi (Sér. Zool.) 12 (1): 3, 4, female, male, nest (in subgenus Monocyttarus) - “Edge of seasonally
flooded evergreen forest, 100 m north of the hamlet of Boca do Mamirauá, which
lies at the eastern end of Mamirauá Ecological Reserve, and 1 km west of the
confluence of the rivers Solimões and Japurá in Amazonas State, Brazil (2°36’
S; 67°45’W)” (holotype female coll. Raw).
Distr.: Brazil: Amazonas.
maracaensis Raw
Mischocyttarus
maracaensis Raw, 1989, Maracá Rainfor. Proj.,
Invert. Limnol., Prelim. Rep.: 30, female, nest (in subgenus Monocyttarus) - “Seasonal evergreen
forest 4 km from eastern end of the Ilha de Maracá, and 1 km north of the
Maracá branch of the river Uraricoera, Territory of Roraima, Brazil” (coll.
Raw); 1989, Maracá Rainfor. Proj., Invert. Limnol., Prelim. Rep.: 41, 42
(habitat preference); 1992, Nature Dynam. For.-Savan. Boundaries: 510; 1998,
Maracá, Biodivers. Environ. Amazon. Rainfor.: 317; 2000 (1999), Rev. Bras.
Zool. 16 (4): 987 (“sp.n.”; described).
Distr.: Brazil: Roraima.
marginatus (Fox)
Polybia
marginata Fox, 1898, Proc. Acad. Nat. Sci. Philad.:
456 - “
Mischocyttarus marginatus; Richards,
1941, Proc. R. Ent. Soc. London (B) 10: 126 (in subgenus Kappa, flavitarsis
group); 1945, Trans. R. Ent. Soc. London 95: 315 (key), 368, fig. 58 (in
subgenus Kappa, flavitarsis group). - Zikán, 1949, Bol. Parq. Nac. Itatiaia 1: 184,
figs. 123, 272. - Richards, 1978, Soc. Wasps Am.: 310 (key), 326 (in subgenus Monocyttarus). - Almeida and Castro,
1991, Acta Biol. Par.,
Distr.: Brazil: D. F., Goiás, Mato Grosso, Minas Gerais,
São Paulo, Rio de Janeiro; Paraguay.
mexicanus cubicola Richards
Mischocyttarus
cubensis; Bequaert, 1933, Entomologica Am. 13: 115 (key),
118 (nest), 135, fig. 4, pl. XXIX figs. 5-6. - Richards, 1941, Proc. R. Ent.
Soc. London (B) 10: 126 (in subgenus Kappa,
cubensis group); 1945, Trans. R. Ent.
Soc. London 95: 317 (key), 381, fig. 69 (in subgenus Kappa, cubensis group). -
Zikán, 1949, Bol. Parq. Nac. Itatiaia 1: 172, figs. 108-110, 252-254. Misidentification.
Mischocyttarus
cubensis var. cubensis;
Bequaert, 1933, Ent. Am. (N. S.) 13: 138 (key), 139 (nest), 142 (
Mischocyttarus
cubensis cubensis; Bohart, 1951, in Muesebeck, et al., Hym. Am. N. of
Mischocyttarus mexicanus; Litte, 1977,
Behav. Ecol. Sociobiol. 2: 229-246 (ethology). - Post and Jeanne, 1982, J.
Mischocyttarus
mexicanus cubicola Richards, 1978, Soc. Wasps Am.:
309 (key), 318, female, male (in subgenus Monocyttarus)
- “
Distr.: U. S. A.: SC, GA, FL, AL;
mexicanus maculipes Richards
Mischocyttarus
cubensis var. cubensis;
Bequaert, 1933, Ent. Am. (N.
S.) 13: 142 [partim].
Mischocyttarus mexicanus var. maculipes
Richards, 1945, Trans. R. Ent. Soc. London 95: 317
(key), 383, female (in subgenus Kappa,
cubensis group) - “
Distr.:
mexicanus mexicanus (de Saussure)
Polybia
mexicana de Saussure, 1854, Ét. Fam. Vesp. 2: 203,
female (in division My) - “Le
Mexique” (lectotype
Megacanthopus indeterminabilis var. mexicana; Ducke, 1910, Ann. Hist.-Nat. Mus. Natl. Hung. 8: 540.
Mischocyttarus cubensis var.
mexicanus; Bequaert, 1933, Ent. Am.
(N. S.) 13: 138 (key), 142 (Guatemala; Honduras); 1940, Bull. Brooklyn Entomol.
Soc. 35: 140 (introduced U. S. A.: TX); 1942; Bull.
Mischocyttarus
cubensis var. sternalis
Richards, 1945, Trans. R. Ent. Soc. London 95: 317 (key), 385, female, male
(in subgenus Kappa, cubensis group) - “
Mischocyttarus
cubensis var.
flavissimus Richards, 1945, Trans. R. Ent. Soc. London 95: 317 (key), 386,
female (in subgenus Kappa, cubensis group) - [
Mischocyttarus mexicanus; Zikán, 1949,
Bol. Parq. Nac. Itatiaia 1: 165, figs. 103, 247. - Strassmann et al., 1995, Ins. Soc. 42 (4): 379-383
(genetic relatedness). - West-Eberhard et
al., 1995, Hym.
Mischocyttarus
cubensis mexicanus; Bohart, 1951, in Muesebeck, et al., Hym. Am. N. of
Mischocyttarus mexicanus morph flavissimus; Richards, 1978, Soc. Wasps
Am.: 308 (key).
Mischocyttarus mexicanus morph sternalis; Richards, 1978, Soc. Wasps
Am.: 308 (key).
Mischocyttarus mexicanus mexicanus; Richards, 1978, Soc. Wasps
Am.: 308 (key), 318 (in subgenus Monocyttarus).
- Rodríguez-Palafox, 1988, Fol. Ent. Mex. 77: 500 (key), 506, fig. 4.1, 4.3
(Mexico: Jalisco); 1996, in Llorente-Bousquets et al., Biodiv. Taxon.
Biogeogr. Artrópod. México: 482 (list).
Distr.: U. S. A.: TX; Mexico: Guerrero,
Jalisco, Michoacán, Morelos, Oaxaca, Puebla, Tamaulipas, Veracruz, Yucatán;
Guatemala; Honduras; Costa Rica.
mimicus Zikán
Mischocyttarus
mimicus Zikán, 1935, Arq. Inst. Biol. Veget. 1 (3):
154 (key), 189, figs. 57, 60, pl. 3 fig. 2, female, nest - [Brazil] “Südabhange
des Itatiaya in etwa 900 m Seehöhe” (Inst. Oswaldo Cruz, Rio de Janeiro); 1949,
Bol. Parq. Nac. Itatiaia 1: 142. - Richards, 1978, Soc. Wasps Am.: 315 (key),
365 (in subgenus Monocyttarus).
Distr.: Brazil: Rio de Janeiro.
mixtus Richards
Mischocyttarus
mixtus Richards, 1978, Soc. Wasps Am.: 309 (key),
319, female (in subgenus Monocyttarus)
- “PANAMA: Bugaba, 800-1500 ft” (London).
Distr.: Panama.
mourei Zikán
Mischocyttarus
mourei Zikán, 1949, Bol. Parq. Nac. Itatiaia
1: 126, figs. 72-73, 211-212, 314, 373, female, male - [Brazil] “Curitiba,
Paraná” (lectotype male Inst. Oswaldo Cruz, Rio de Janeiro); also from São
Paulo; Santa Catarina. - Richards, 1978, Soc. Wasps Am.: 313
(key), 346 (designation of lectotype; in subgenus Monocyttarus).
Distr.: Brazil: Paraná, Santa Catarina,
São Paulo.
mutator Zikán
Mischocyttarus
mutator Zikán, 1949, Bol. Parq. Nac. Itatiaia 1:
178, fig. 262, female - [Brazil] “Itatiaia, 550 m (Fazenda ‘Valparaiso’)”
(lectotype Inst. Oswaldo Cruz, Rio de Janeiro). - Richards, 1978, Soc. Wasps
Am.: 314 (key), 361 (designation of lectotype; in subgenus Monocyttarus).
Distr.: Brazil: Rio de Janeiro.
navajo Bequaert
Polybia
flavitarsis; Viereck, 1906, Trans. Am. Ent. Soc.
32: 231 (
Megacanthopus flavitarsis; Pierce,
1918, Proc.
Mischocyttarus flavitarsis; Salt and
Bequaert, 1929, Psyche 36: 261. - Litte, 1979, Z. Tierpsychol. 50: 282-312
(ethology).
Mischocyttarus
flavitarsis var. navajo Bequaert, 1933, Ent. Am. (N. S.) 13: 122 (key), 125, 126
(nest), 127 (stylopization), 131, pl. XXIX figs. 1-2, female, male, nest - [
Mischocyttarus flavitarsis navajo; Bohart, 1951, in Muesebeck, et al., Hym. Am. N. of
Mischocyttarus
navajo; Snelling, 1983, Pan-Pacif. Ent. 59: 277. -
Rodríguez-Palafox, 1996, in Llorente-Bousquets et al., Biodiv.
Taxon. Biogeogr. Artrópod. México: 482 (list).
Distr.:
oreophilus Zikán
Mischocyttarus
“creophilus
Zikán nov. spec.” Weyrauch,
1943, Bol. Mus. Hist. Nat. Javier Prado 7: 68. Nomen
nudum.
Mischocyttarus oreophilus Zikán, 1949, Bol. Parq. Nac. Itatiaia 1: 134, female - “Machupichu,
2.000 m (Peru)” (lectotype Inst. Oswaldo Cruz, Rio de
Janeiro). - Richards, 1978, Soc. Wasps Am.: 309 (key), 325 (designation of
lectotype; in subgenus Monocyttarus).
Distr.: Colombia; Peru: Cuzco.
pallidipectus (Smith)
Polybia pallidi-pectus Smith, 1857, Cat.
Hym. Ins. Coll. Brit. Mus. 5, Vesp.: 128, male - “Mexico” (London). - Dalla
Torre, 1894, Cat. Hym. 9: 165 (pallidipecta;
cat.); 1904, Gen. Ins. 19: 78 (cat.). - Ducke, 1910, Ann. Hist.-Nat. Mus. Natl. Hung. 8: 541 (syn.
of M. rufidens (de Saussure)).
Mischocyttarus pallidipectus;
Richards, 1941, Proc. R. Ent. Soc. London (B) 10: 126 (in subgenus Kappa, flavitarsis group); 1945, Trans. R. Ent. Soc. London 95: 314 (key),
361, figs. 45-46, pl. 1 fig. 123 (in subgenus Kappa, flavitarsis
group). - Zikán, 1949, Bol. Parq. Nac. Itatiaia 1: 183,
figs. 121-122, 269-271, 299, 317-318. - Snelling, 1970, Contrib. Sci. 137: 18
(compared to fisheri n. sp.). -
Richards, 1978, Soc. Wasps Am.: 309 (key), 324 (syn.: M. silvaticus Zikán; in subgenus Monocyttarus). - West-Eberhard et
al., 1995, Hym. Costa Rica: 585 (Costa Rica). -
Rodríguez-Palafox, 1996, in Llorente-Bousquets et al., Biodiv. Taxon.
Biogeogr. Artrópod. México: 482 (list).
Mischocyttarus
oaxacanus Zikán, 1949, Bol. Parq. Nac. Itatiaia 1: 32
(key), 168, figs. 105-106, 249-250, female, male - [Mexico:] “Basse Mistéque,
Env. d’Oaxaca...México...Galeana, Nueva Leon (3-6.000 pés)” (São Paulo). -
Snelling, 1983, Pan-Pacif. Ent. 59: 275 (syn. of M. immarginatus Richards). - Silveira,
2006, Zootaxa 1321: 23 (syn. of M. pallidipectus (Smith)).
Mischocyttarus silvaticus Zikán, 1949,
Bol. Parq. Nac. Itatiaia 1: 190, figs. 126, 276, 297, 410,
female, male - “San José da Costa Rica” (lectotype female Inst. Oswaldo Cruz, Rio de Janeiro). - Richards, 1978, Soc. Wasps Am.: 324
(designation of lectotype; syn. of M.
hirtulus Zikán).
Distr.: Mexico: Chiapas, Guerrero, Morelos,
Oaxaca, Puebla, Veracruz; Guatemala; Honduras; Costa Rica; Panama.
pallidipes Richards
Mischocyttarus
pallidipes Richards, 1945, Trans. R. Ent. Soc.
London 95: 315 (key), 366, female (in subgenus Kappa, flavitarsis group)
- “BRAZIL, Santa Catharina, Thersepolis” [Theresepolis] (Berlin); 1978, Soc.
Wasps Am.: 362.
Distr.: Brazil: Santa Catarina.
paraguayensis Zikán
Mischocyttarus
paraguayensis Zikán, 1935, Arq. Inst. Biol. Veget.
1 (3): 154 (key), 165, figs. 23, 25, 28, 30, pl. 3 fig. 3, female, male, nest -
“Mborero in
Mischocyttarus alfkenii var. zikanii Richards, 1945, Trans. R. Ent. Soc. London 95: 318 (key),
399, female, male (in subgenus Kappa,
cubensis group) - “PARAGUAY: Alto
Parana, Puerto Bertoni” (holotype female London); also from four other
localities.
Distr.: Brazil: Rio Grande do Sul, Santa Catarina;
Paraguay: Guairá, Alto Paraná, Asunción; Argentina: Jujuy, Salta, Tucumán,
Misiones.
peduncularius Zikán
Mischocyttarus “peduncularius Zikán nov. spec.” Weyrauch, 1943, Bol. Mus. Hist. Nat.
Javier Prado 7: 68. Nomen nudum.
Mischocyttarus peduncularius Zikán, 1949, Bol. Parq. Nac. Itatiaia
1: 151, figs. 232-234, 294, female, male - “Peru (Vale Chanchamayo, 800 a 1.200
metros)” (lectotype male Inst. Oswaldo Cruz,
Distr.: Peru: Junín.
petiolatus Richards
Mischocyttarus
petiolatus Richards, 1978, Soc. Wasps Am.: 312
(key), 336, female, male (in subgenus Monocyttarus)
- “PERU: Piura, Jaen, 700 m” (holotype female London); also from Cajamarca; and
Colombia. - Sarmiento, 1994, Rev. Biol. Trop. 42 (1/2): 361 (list).
Distr.: Colombia: Putumayo; Peru:
Cajamarca, Piura.
phthisicus (Fabricius)
Vespa
phthisica Fabricius, 1793, Ent. Syst. 2: 281 - “America” (Kobenhavn). -
Ducke, 1910, Ann. Hist.-Nat. Mus. Natl. Hung. 8: 541 (? Polybia, Stelopolybia or Megacanthopus).
Polistes phthisica; Fabricius, 1804, Syst. Piez.: 278.
Polybia indeterminabilis de Saussure, 1854, Ét. Fam. Vesp. 2: 201 (in division My) - “Amérique du Sud. Ile
Sainte-Lucie” (lectotype female Paris). - Dalla Torre, 1894, Cat. Hym. 9: 164
(cat.); 1904, Gen. Ins. 19: 77 (cat.). - Ducke, 1913, Deutsch. Ent. Zeitschr.: 331 (syn. of M.
phthisicus (Fabricius, 1793)). - Bequaert, 1933, Entomologica Am. 13: 116
(identity uncertain). - Richards, 1978, Soc. Wasps Am.: 316 (designation of
lectotype). - Carpenter, 1999, Am. Mus. Novitat. 3259: 21 (notes on types).
Polybia
mexicana; Ashmead, 1900, Trans. Ent. Soc. Lond. 311
(Puerto Rico). Misidentification.
Megacanthopus indeterminabilis; Ducke,
1910, Ann. Hist.-Nat. Mus. Natl. Hung. 8: 540.
Mischocyttarus phthisicus; Ducke, 1913,
Deutsch. Ent. Zeitschr.: 331 (syns.: P.
indeterminabilis de Saussure, cubensis
de Saussure, mexicanus de Saussure);
1914, Zool. Jahrb. (Syst.) 36: 328. - da Fonseca, 1926, Rev. Mus. Paul. 14: 181
(compared to ypiranguensis n. sp.),
pl. 2 fig. 3 (nest). - Bequaert, 1933, Ent. Am. (N. S.) 13: 116 (key), 139 (St.
Croix). - Richards, 1941, Proc. R. Ent. Soc. London (B) 10: 126 (in subgenus Kappa, cubensis group); 1945, Trans. R. Ent. Soc. London 95: 316 (key),
400 (in subgenus Kappa, cubensis group). - Zikán, 1949, Bol.
Parq. Nac. Itatiaia 1: 171, fig. 107. - Richards, 1978, Soc. Wasps Am.: 308
(key), 316 (in subgenus Monocyttarus).
- Sarmiento, 1994, Rev. Biol. Trop. 42 (1/2): 360 (list).
Mischocyttarus indeterminabilis; Zikán,
1949, Bol. Parq. Nac. Itatiaia 1: 212. - Overal, 1978, Bol. Mus. Para. Emílio
Goeldi (N. S.), Zool. 94: 10 (list).
Distr.: U. S. A.: FL; Haiti; Dominican Republic; Puerto
Rico; Virgin Is.; St. Lucia; Guatemala; Colombia (?).
piceus Zikán
? Mischocyttarus “fusconiger Zikán nov. spec.” Weyrauch, 1943, Bol. Mus. Hist. Nat.
Javier Prado 7: 68. Nomen nudum.
Mischocyttarus piceus Zikán, 1949, Bol. Parq. Nac. Itatiaia 1: 137, fig.
347, female - [Peru] “Oxapampa, 1.700-1.800 m,...Vale Chanchamayo, 1.400 m” (? Inst.
Oswaldo Cruz, Rio de Janeiro).
Distr.: Peru:
Junín.
plaumanni
Zikán
Mischocyttarus plaumanni Zikán, 1949, Bol. Parq. Nac. Itatiaia 1:
167, figs. 104, 383, female - [Brazil] “Nova Teutônia, sul de Santa Catarina”
(lectotype Inst. Oswaldo Cruz, Rio de Janeiro). - Richards,
1978, Soc. Wasps Am.: 312 (key), 333 (in subgenus Monocyttarus).
Distr.: Brazil: Paraná (?), Santa Catarina, São Paulo
(?).
proximus Zikán
Mischocyttarus
proximus Zikán, 1949, Bol. Parq. Nac. Itatiaia 1: 26 (key), 130, female
[also listed as male] - “Campos da Serra, Estado de S. Paulo, Brasil” (São
Paulo). - Richards, 1978, Soc. Wasps Am.: 312 (key), 339 (in
subgenus Monocyttarus).
Distr.: Brazil: São Paulo.
riograndensis Richards
Mischocyttarus
riograndensis Richards, 1978, Soc. Wasps Am.: 314
(key), 361, female, male (in subgenus Monocyttarus)
- “BRAZIL: RS, S. Leopoldo” (holotype female São Paulo); also from Nova
Teutônia. - Carpenter, 1999, Am. Mus. Novitat. 3259: 22 (note on type).
Distr.: Brazil: Rio Grande do Sul.
rivulorum Richards
Mischocyttarus
rivulorum Richards, 1978, Soc. Wasps Am.: 314
(key), 354, female, male (in subgenus Monocyttarus)
- “ARGENTINA: Entre Rios, Rio Uruguay, Isla Pepe-Aji” (holotype female
Tucumán).
Distr.:
Argentina: Entre Rios, Misiones.
rufidens (de Saussure)
Polybia rufidens de Saussure, 1854, Ét. Fam.
Vesp. 2: 204, female (in division My)
- [French Guiana] “Cayenne” (type depository unknown). - Dalla Torre, 1894,
Cat. Hym. 9: 166 (cat.); 1904, Gen. Ins. 19: 78 (cat.). - Richards, 1978, Soc.
Wasps Am.: 318. - Carpenter, 1999, Am. Mus. Novitat. 3259: 22.
Megacanthopus
rufidens; Ducke, 1908, Rev. Ent., Caen 27: 86
(Brazil); 1910, Ann. Hist.-Nat.
Mus. Natl. Hung. 8: 541.
Mischocyttarus rufidens;
Ducke, 1918, Rev. Mus. Paul. 10: 352. - Richards, 1940,
Ann. Mag. Nat. Hist. (11) 5: 183 (“may be the same as M. duckei (du Buysson, 1908)”); 1945, Trans. R. Ent. Soc. London
95: 385 (“there is not even complete certainty as to its correct generic
position”). - Zikán, 1949, Bol. Parq. Nac. Itatiaia 1: 212. - Overal,
1978, Bol. Mus. Para. Emílio Goeldi (N. S.), Zool. 94: 10 (list).
Distr.: French Guiana.
rufipes Zikán
Mischocyttarus
rufipes Zikán, 1949, Bol. Parq. Nac. Itatiaia 1: 135, female -
“Huacapistana, 1.800 m, (Peru)” (Inst. Oswaldo Cruz,
Distr.:
rufomaculatus Richards
Mischocyttarus
hirsutus var. rufomaculatus
Richards, 1945, Trans. R. Ent. Soc. London 95: 315 (key), 364, female (in
subgenus Kappa, flavitarsis group) - “
Mischocyttarus rufomaculatus; Richards,
1978, Soc. Wasps Am.: 310 (key), 327 (in subgenus Monocyttarus). - Sarmiento, 1994, Rev. Biol. Trop. 42 (1/2):
360 (Colombia: Huila, Putumayo; list).
Distr.: Colombia: Huila, Putumayo; Ecuador: Tungurahua;
Bolivia: La Paz.
scitulus Zikán
Mischocyttarus
scitulus Zikán, 1949, Bol. Parq. Nac. Itatiaia 1: 176, figs. 113, 258-259,
female, male - [Brazil] “Itatiaia, 700 m, Est. do Rio de Janeiro” (lectotype
male Inst. Oswaldo Cruz, Rio de Janeiro). - Richards, 1978, Soc. Wasps Am.: 314 (key), 359 (designation of lectotype; in subgenus Monocyttarus).
Distr.: Brazil: Rio de Janeiro.
similaris Zikán
Mischocyttarus
similaris Zikán, 1949, Bol. Parq. Nac. Itatiaia 1:
167, fig. 248, female - [Brazil] “Ilha Sêca, Estado de São Paulo” (Inst.
Oswaldo Cruz, Rio de Janeiro). - Richards, 1978, Soc. Wasps Am.: 314 (key), 360
(in subgenus Monocyttarus).
Distr.: Brazil: São Paulo.
similatus Zikán
Mischocyttarus similatus Zikán, 1935, Arq. Inst. Biol. Veget. 1 (3): 154
(key), 187, figs. 51, 53, pl. 3 fig. 7, female, nest - [Brazil] “Gelände der
Fazenda ‘Valparaizo’ in etwa 550 m Seehöhe” (Inst. Oswaldo Cruz, Rio de
Janeiro); 1949, Bol. Parq. Nac. Itatiaia 1: 138. -
Richards, 1978, Soc. Wasps Am.: 314 (key), 357 (in subgenus Monocyttarus).
Distr.: Brazil: Rio de Janeiro.
tarmensis Richards
Mischocyttarus
hirsutus var. tarmensis Richards,
1945, Trans. R. Ent. Soc. London 95: 315 (key), 365, female, male (in subgenus Kappa, flavitarsis group) - “PERU: Rio Tarma, Huacapistana” (holotype
female Ithaca); also from another locality.
Mischocyttarus “huacapistanus Zikán nov. spec.” Weyrauch, 1943, Bol. Mus. Hist. Nat.
Javier Prado 7: 69. Nomen nudum.
Mischocyttarus huacapistanus Zikán, 1949, Bol. Parq. Nac. Itatiaia 1:
133, 236 (“host” of huacapistanoides
n. sp.), figs. 82, 221-222, female, male - “Huacapistana, 1.800 m, ... Peru”
(lectotype male Inst. Oswaldo Cruz, Rio de Janeiro); also from Oreja de Capelo.
- Richards, 1978, Soc. Wasps Am.: 332 (designation of
lectotype; syn. of M. tarmensis Richards).
Mischocyttarus
huacapistanoides Zikán, 1949, Bol. Parq. Nac.
Itatiaia 1: 236, fig. 351, locality not stated (lectotype male Inst. Oswaldo
Cruz,
Mischocyttarus tarmensis; Richards,
1978, Soc. Wasps Am.: 310 (key), 332 (syns.: M. huacapistanus Zikán, M.
huacapistanoides Zikán; in subgenus Monocyttarus).
Distr.:
transandinus Richards
Mischocyttarus
transandinus Richards, 1978, Soc. Wasps Am.: 312
(key), 338, female, male (in subgenus Monocyttarus)
- “
Distr.: Peru:
Lambayeque.
wagneri
(du Buysson)
Megacanthopus Wagneri du Buysson, 1908, Rev. Ent., Caen 27: 219, female
- “Brésil, dans les montagnes des Orgues, aux environs de la Tijuca, par 1.000
mètres d’altitude” (Paris).
Mischocyttarus
pedunculatus Zikán, 1935, Arq. Inst. Biol. Veget. 1
(3): 155 (key), 159, figs. 9-10, 14, pl. 2 fig. 6, pl. 3 fig. 11, female, male,
nest - [Brazil] “Südabhange des Itatiaya in 500 m. (Fazenda Valparaiso)”
(lectotype female São Paulo); also from 700 m. - Richards, 1941, Proc. R. Ent.
Soc. London (B) 10: 126 (syn. of M.
wagneri (du Buysson)).- W. Zikán and Wydodzinsky, 1948, Bol. Serv. Nac.
Pesq. Agron. 4: 75 (list). - Zikán, 1949, Bol. Parq. Nac. Itatiaia 1: 149, 236
(“host” of pedunculatoides n. sp.),
figs. 92-93, 232, 327, 381, 418-419. - Richards, 1978, Soc. Wasps Am.: 334
(designation of lectotype; syn.: M.
pedunculatoides Zikán). - Carpenter, 1999, Am. Mus. Novitat. 3259: 22
(notes on types).
Mischocyttarus wagneri; Richards, 1941,
Proc. R. Ent. Soc. London (B) 10: 126 (in subgenus Kappa, flavitarsis group;
syn.: M. pedunculatus Zikán); 1945,
Trans. R. Ent. Soc. London 95: 315 (key), 371, fig. 50, pl. 1 fig. 122 (in
subgenus Kappa, flavitarsis group). - Zikán, 1949, Bol. Parq. Nac. Itatiaia 1: 150.
- Richards, 1978, Soc. Wasps Am.: 312 (key), 334, fig. 114 (in subgenus Monocyttarus).
Mischocyttarus
pedunculatoides Zikán, 1949, Bol. Parq. Nac.
Itatiaia 1: 236, sex and locality not stated (lectotype female Inst. Oswaldo
Cruz,
Distr.: Brazil: Minas Gerais, Rio de
Janeiro, São Paulo.
Subgenus
Scytokeraia Cooper
Scytokeraia
Cooper, 1997, Ent. Mon.
Mag. 133: 117, subgenus of Mischocyttarus
de Saussure.
Type species: Mischocyttarus mastigophorus
Richards, 1978, by original designation.
alienus
Richards
Mischocyttarus alienus Richards, 1978, Soc. Wasps Am.: 375 (key), 378,
female (in subgenus Monogynoecus) -
“COLOMBIA: Valle, Anchicayá nr Buenaventura, 300 m” (London). - Carpenter and
Wenzel, 1988, Psyche 95: 90-92, 96, fig. 10 (Boliva; Costa Rica). - Sarmiento,
1994, Rev. Biol. Trop. 42 (1/2): 361 (list). - West-Eberhard et al., 1995, Hym. Costa Rica: 585
(Costa Rica). - Cooper, 1996, Ent. Mon. Mag. 132: 23 (removed from Monogynoecus) ; 1997, Ent. Mon. Mag.
133: 117, 118 (key), 124 (in subgenus Scytokeraia;
male, nest; distr.: Ecuador), figs. 1-2, 9-11.
Distr.: Costa Rica; Colombia: Valle; Ecuador: Pichincha.
fraudulentus Richards
Mischocyttarus
fraudulentus Richards, 1978, Soc. Wasps Am.: 375 (key), 379, female (in
subgenus Monogynoecus) - “COLOMBIA:
Nariño, Barbacoas, 80 m” (London). - Carpenter and Wenzel, 1988, Psyche 95:
90-92, 94, figs. 2, 5, 9 (Costa Rica). - Sarmiento, 1994, Rev. Biol. Trop. 42
(1/2): 361 (list). - West-Eberhard et al.,
1995, Hym. Costa Rica: 585 (Costa Rica). - Cooper, 1996, Ent. Mon. Mag. 132: 23
(removed from Monogynoecus); 1997,
Ent. Mon. Mag. 133: 117, 118 (key, in subgenus Scytokeraia).
Distr.: Costa Rica; Colombia: Nariño.
juditai Cooper
Mischocyttarus
juditai Cooper, 1997, Ent. Mon. Mag. 133: 117, 119 (key), 122, female,
male, nest, figs. 5, 12 (in subgenus Scytokeraia)
- “ECUADOR: Morona-Santiago, Macas, 1,1000 m” (holotype male London); also from
Pastaza; Colombia.
Distr.: Ecuador: Pastaza;Colombia: Caqueta.
mastigophorus Richards
Mischocyttarus
mastigophorus Richards, 1978, Soc. Wasps Am.: 375
(key), 376, male not female (in subgenus Monogynoecus)
- “ECUADOR: [El Oro], near Piñas, Morro Morro” (holotype male New York); also
from Mexico. - West-Eberhard et al.,
1995, Hym. Costa Rica: 585 (Costa Rica). - Cooper, 1996, Ent. Mon. Mag. 132: 23
(removed from Monogynoecus); 1997,
Ent. Mon. Mag. 133: 117 (type of new subgenus Scytokeraia), 118 (key), 124 (female; distr.: Costa Rica), fig. 3.
- O’Donnell, 1998, Behav. Ecol. Sociobiol. 43: 327-331 (ethology). - Carpenter,
1999, Am. Mus. Novitat. 3259: 21 (notes on types). - O’Donnell, 1999, Ethology
105: 273-282 (ethology). - O’Donnell and Joyce, 1999, Biol J. Linn. Soc. 66:
501-514 (mimicry of Agelaia xanthopus
and A. yepocapa).
Distr.: Costa Rica; Ecuador: El Oro.
moralesi Zikán
Mischocyttarus
moralesi Weyrauch, 1943, Bol. Mus. Hist. Nat. Javier Prado 7: 69. Nomen nudum.
Mischocyttarus moralesi Zikán, 1949, Bol. Parq. Nac. Itatiaia 1: 146, fig.
229, female - “Vale de Chanchamayo, 800 m, (Peru)” (Inst. Oswaldo Cruz, Rio de
Janeiro). - Richards, 1978, Soc. Wasps Am.: 375 (key), 379 (in subgenus Monogynoecus). - Carpenter and Wenzel,
1988, Psyche 95: 92. - Cooper, 1996, Ent. Mon. Mag. 132: 23 (removed from Monogynoecus); 1997, Ent. Mon. Mag. 133:
117, 118 (key), 126 (in subgenus Scytokeraia;
nest; distr.).
Distr.: Ecuador: Morona-Santiago; Peru: Junín; Bolivia:
La Paz.
pelor Carpenter
Mischocyttarus
pelor Carpenter, 1988, in
Carpenter and Wenzel, Psyche 95: 89, figs. 1, 3, 6, 8, female, nest (in
subgenus Monogynoecus) - “Turrialba,
Costa Rica” (Washington). - West-Eberhard et
al., 1995, Hym. Costa Rica: 585 (Costa Rica). - Cooper,
1996, Ent. Mon. Mag. 132: 23 (removed from Monogynoecus)
; 1997, Ent. Mon. Mag. 133: 117, 118 (key, in subgenus Scytokeraia).
Distr.: Costa Rica.
rhadinomerus Cooper
Mischocyttarus
rhadinomerus Cooper 1997, Ent. Mon. Mag. 133: 117,
118 (key), 121, female, nest, fig. 7 (in subgenus Scytokeraia) - “PERU: Amazonas, 20 km S.E. Rodriguez de Mendoza,
1,1800 m” (London).
Distr.: Peru: Amazonas.
sprucei Cooper
Mischocyttarus
sprucei Cooper 1997, Ent. Mon. Mag. 133: 117, 119
(key), 121, female, nest, fig. 8 (in subgenus Scytokeraia) - “ECUADOR: Tungurahua, Pastaza Valley, Rio Topo, 1,1000
m” (London); also from Bolivia.
Distr.: Ecuador: Pastaza; Bolivia: La Paz.
stenoecus Richards
Mischocyttarus
stenoecus Richards, 1978, Soc. Wasps Am.: 390
(key), 399, female (in subgenus Haplometrobius)
- “COLOMBIA: Nariño, Barbacoas, 50 m” (London). - Sarmiento, 1994, Rev. Biol.
Trop. 42 (1/2): 361 (list). - Cooper, 1997, Ent. Mon. Mag. 133: 117, 118 (key),
128 (in subgenus Scytokeraia; male,
nest; distr.: Colombia: Valle), figs. 6, 13.
Distr.: Colombia: Nariño, Valle.
subornatus Zikán
Mischocyttarus
“modestus Zikán
nov. spec.” Weyrauch, 1943,
Bol. Mus. Hist. Nat. Javier Prado 7: 69.
Nomen nudum.
Mischocyttarus subornatus Zikán, 1949, Bol. Parq. Nac. Itatiaia 1:
124, figs. 320, 372, female - “Oxapampa, 1.800 m, Peru” (lectotype Inst. Oswaldo
Cruz, Rio de Janeiro). - Richards, 1978, Soc. Wasps Am.: 308 (key), 316
(designation of lectotype; in subgenus Monocyttarus).
- Cooper, 1997, Ent. Mon. Mag. 133: 117, 118 (key), 125 (in subgenus Scytokeraia; male, nest; distr.: Peru:
Junín), fig. 4.
Distr.: Peru: Pasco, Junín
telembi Cooper
Mischocyttarus
telembi Cooper 1997, Ent. Mon. Mag. 133: 117, 118 (key), 120, female, nest
(in subgenus Scytokeraia) -
“COLOMBIA: Nariño, Barbacoas (50 m)” (holotype male London); also from Pastaza;
Colombia.
Distr.: Colombia: Nariño.
veracrucis Cooper
Mischocyttarus
veracrucis Cooper 1997, Ent. Mon. Mag. 133: 117, 118 (key), 119, female (in
subgenus Scytokeraia) - “MEXICO: V.C.
nr Huatusco” (holotype Washington); also from Pastaza; Colombia.
Distr.:
Unrecognized
Species
“Megacanthopus melenopygoides Weyrauch” Colomo and Berta,
2005, Rev. Soc. Entomol. Argent. 64: 83. Nomen
nudum.
“Mischocyttarus gentili Zikán” Colomo
and Berta, 2005, Rev. Soc. Entomol. Argent. 64: 83. Nomen
nudum.
Mischocyttarus “parallelus Zikán nov. spec.” Weyrauch, 1943, Bol. Mus. Hist. Nat. Javier
Prado 7: 69. Nomen nudum.
“Mischocyttarus subventris Sauss.” Overal, 1978, Bol. Mus. Para. Emílio Goeldi (N.
S.), Zool. 94: 10. Nomen nudum.
Mischocyttarus undulatus; Richards, 1945, Trans. R.
Ent. Soc. London 95: 338; 1978, Soc. Wasps Am.: 393 (key), 413 (in subgenus Haplometrobius; Brazil) [misidentification]. - Sarmiento, 1994, Rev. Biol. Trop.
42 (1/2): 362 (Colombia: Putumayo; list). - Mejdalani and Felix, 1997, Proc.
Entomol. Soc. Wash. 99 (1):159 (mimicry by Lissocarta
beckeri (Homoptera: Cicadellidae)). - Carpenter, 1999, Am. Mus. Novitat. 3259: 23 (undulatus
misidentified by Richards).
Polybia carbonaria ?; Fox, 1898,
Proc. Acad. Nat. Sci. Philad.: 450, male -
* indicates new record.