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INTRODUCTION
Snakes of the species rich genus Lycodon Boie, 1826 re-
ceived considerable attention in most regions of Asia. Six
new species were described form the Philippines (Ota &
Ross 1994; Lanza 1999; Gaulke 2002) and new species
were discovered in Cambodia (Daltry & Wüster 2002), In-
dia (Mukherjee & Bhupathy 2007) and Myanmar (Slowin-
ski et al., 2001). The taxonomy of the Chinese part of the
genus remained unattended until recently. Pope (1935) list-
ed five species, namely Lycodon capucinus Boie, 1827,
Lycodon fasciatus (Anderson, 1879), Lycodon laoensis
Günther, 1864, Lycodon ruhstrati (Fischer, 1886) and Ly-
codon subcinctus Boie, 1827. This arrangement has not
changed for the next 75 years. Vogel et al. (2010) reviewed
the Lycodon ruhstrati complex and described Lycodon
ruhstrati abditus as a new subspecies from China, and
revalidated Lycodon futsingensis (Pope, 1928). Detailed
examination of banded specimens of the genus led us to
the conclusion, that the diversity is much higher in this
region and that several species await description.
In the course of our ongoing review of the Lycodon fas-
ciatus complex, we came upon two specimens of the genus
Lycodon from Yunnan, China, which seemed to be differ-
ent from L. fasciatus. Adetailed examination showed clear
morphological differences which lead us to describe them
as new species.
MATERIAL & METHODS
This revision is based on a total of 67 preserved specimens
of Lycodon fasciatus auctorum examined for their exter-
nal morphological characters and on several photographed
specimens. They are listed in the Appendix I. Compara-
tive material of the L. ruhstrati complex is listed under
Vogel et al. (2010).
A total of 53 morphological characters were recorded for
each specimen. The characters and their abbreviations are
listed in Table 1. Not all of these characters have been used
for this study, but all of them were compared.
Measurements, except body and tail lengths, were taken
with a slide-caliper to the nearest 0.1 mm; all body meas-
urements were made to the nearest millimetre. The num-
ber of ventral scales was counted according to Dowling
(1951). Half ventrals were not counted except they were
present on both sides (divided ventrals). The terminal scute
is not included in the number of subcaudals. The dorsal
scale row counts are given at one head length behind head,
at midbody (i.e., at the level of the ventral plate correspon-
ding to a half of the total number of ventrals), and at one
head length before vent. We considered being sublabials
those shields that were completely below a supralabial.
Values for paired head characters are given in left / right
order.
A new species of the genus Lycodon (Boie, 1826)
from Yunnan Province, China
(Serpentes: Colubridae)
Gernot Vogel1& Patrick David2
1Society for Southeast Asian Herpetology, Im Sand 3, D-69115 Heidelberg, Germany;
E-mail: Gernot.Vogel@t-online.de
2Muséum national d’Histoire naturelle, Département Systématique et Évolution, Reptiles & Amphibiens,
UMR 7205 OSEB, Case postale 30, 57 rue Cuvier, F-75231 Paris Cedex 05, France; E-mail: pdavid@mnhn.fr
Bonn zoological Bulletin Volume 57 Issue 2 pp. 289–296 Bonn, November 2010
Abstract. A new species of the genus Lycodon is described from Yunnan Province, People’s Republic of China. It dif-
fers from the superficially similar Lycodon fasciatus by the fact that the loreal is not entering orbit, in colouration, and
lower numbers of subcaudals and infralabials. From the Lycodon ruhstrati group it differs by the colouration of the ven-
ter and the dorsal bands. This new species is only known the Chinese province of Yunnan.
Keywords. Oriental Region, China, Colubrinae, Lycodon fasciatus, taxonomy, Lycodon synaptor sp. n.
Bonn zoological Bulletin 57 (2): 289–296 ©ZFMK
The white or light bands on the body and tail were count-
ed on one side. Hardly visible or incomplete bands were
counted as one, bands that were fused were counted as
two. The collar on the neck was not counted and bands
covering the anal shield were added to the bands of the
body.
Museum abbreviations
BMNH: The Natural History Museum, London, UK. –
BNHS: Bombay Natural History Society, Mumbai, India.
– CAS: California Academy of Sciences, San Francisco,
USA. – CIB: Chengdu Institute of Biology, Chengdu,
People’s Republic of China. – FMNH: Field Museum of
Natural History, Chicago, USA. – KIZ: Kunming Insti-
tute of Zoology, Kunming, People’s Republic of China.
– MNHN: Muséum national d’Histoire naturelle, Paris,
France. – NMW: Naturhistorisches Museum Wien, Aus-
tria. – ZFMK: Zoologisches Forschungsmuseum
Alexander Koenig, Bonn, Germany. – ZMB: Zoologis-
ches Museum für Naturkunde der Humboldt-Universität
zu Berlin, Berlin, Germany. – ZSM: Zoologische
Staatssammlung, München, Germany.
RESULTS
Lycodon synaptor sp. n.
Holotype. BMNH 1905.1.30.63 adult female (tail dissect-
ed), from “Tongchuan, Yunnan”, today Dongchuan, 100
km north of Kunming, Yunnan Province, People’s Repub-
lic of China (Figs 1–3). Collected by the J. Graham Ex-
pedition, unknown date.
290 Gernot Vogel & Patrick David
Bonn zoological Bulletin 57 (2): 289–296 ©ZFMK
Table 1. List of morphological characters used in this study and
their abbreviation.
Fig. 1. Dorsal view of preserved holotype of Lycodon synap-
tor sp. n., BMNH 1905.1.30.63 from Dongchuan, 100 km north
of Kunming, Yunnan Province, People’s Republic of China. Pho-
tograph by Gernot Vogel.
N° Abbreviation Characters
Morphometry
1 SVL Snout-vent length (mm)
2 TaL Tail length (mm)
3 TL Total length (mm)
4 Rel TL Relative tail length TaL/TL
Anatomy
5 TEETH Number of upper maxill. teeth (one side)
Scalation
6 DSR Dorsal scale rows
7 ASR Dorsal scale rows at neck
8 MSR Dorsal scale rows at midbody
9 PSR Dorsal scale rows before vent
10 Keel Number of keeled dorsal rows
11 VEN Ventral plates
12 PreVEN Number of preventrals
13 VEN not Ventrals notched or not
14 VEN keel Ventrals keeled
15 SC Subcaudal plates
16 ANA Anal plate: 1: single – 2: divided
17 Lor-l Number of loreal scale (0 or 1) at left
18 Lor-r Number of loreal scale (0 or 1) at right
19 Lo touch-l Loreal scale touches eye at left
20 Lo touch-r Loreal scale touches eye at right
21 SL-l Number of supralabials at left
22 SL-r Number of supralabials at right
23 SL/Eye-l Numbers of the SL entering orbit at left
24 SL/Eye-r Numbers of the SL entering orbit at right
25 Larg SL-l Largest SL left
26 Larg SLrl Largest SL right
27 IL-l Number of infralabials at left
28 IL-r Number of infralabials at right
29 IL-tot Total number of infralabials
30 IL/1st child Number of IL in contact with
anterior chin shield
31 PreOc-l Number of preoculars at left
32 PreOc-r Number of preoculars at right
33 PostOc-l Number of postoculars at left
34 PostOc-r Number of postoculars at right
35 ATem-l Number of anterior temporals at left
36 ATem-r Number of anterior temporals at right
37 PTem-l Number of posterior temporals at left
38 PTem-r Number of posterior temporals at right
39 ParaR Temporal row containing paraparietals
40 Paras Plates surrounding paraparietals,
see Inger & Marx (1965)
41 Parab Scales between the paraparietals
Pattern
42 BODCOL Body colour1: grey; 2: brown or ochre
43 Bands Number of bands on body
44 Tail bands Number of bands on tail
45 Tail venter Colouration of tail venter
46 Bellycol Colouration of belly
47 Bellyspeck Speckling of belly
48 First band Number of VEN before the
first band starts, counted left side
49 Broad base Number of VEN that are covered at
the base of the first band
50 Broad vert Numbers of vertebral scales that are
covered by the first band
51 Edged Dorsal bands with light margins
52 Coul throat Colour of the throat
53 Ve throat Dark VEN on the throat before
the first band
Paratype. MNHN 1905.0283, adult female (tail dissect-
ed), from “Tongchuan Fu, Chine”, at present Dongchuan,
Yunnan Province, People’s Republic of China. Collected
by W. F. H. Rosenberg on 21st July 1905.
Diagnosis. A species of the genus Lycodon characterized
by: (1) loreal scale not entering orbit; (2) 15–17 dorsal
scale rows at the forepart of the body and 17 dorsal scale
rows at midbody; (3) upper and vertebral dorsal rows
(6–7) keeled; (4) 201–203 ventrals in females, males un-
known; (5) 68–69 Sc in females, males unknown; (6) a
relative tail length of about 0.189–0.192 in females, males
unknown; (7) 8 supralabials with SL 4–6 touching the or-
bit (7) 30–31 narrow white bands on a dark body; (8)
width of the first band vertebral 0.5–1.0 scales, on the base
3 ventrals; and (9) the belly with discreet bands through-
out.
The new species can be recognized by the combination
of the loreal scale not entering orbit (entering in L. fas-
ciatus sensu stricto), its narrow dorsal bands, with the first
band starting at ventral 5–9 (more irregular in L. fascia-
tus [Fig. 4] and species of the L. ruhstrati group, where
they usually start later) and the dark throat, which usual-
ly is light in other species of the L. fasciatus group and
the L. ruhstrati group. Most other characters match with
Lycodon fasciatus.
Detailed comparisons with other species of the genus Ly-
codon appear below in the Discussion.
Etymology. This species is indirectly named in honour of
Dr. Wolfgang Böhme. It was always a publicized aim of
Wolfgang Böhme to unite professional and amateur her-
petologists. We, both authors have always been amateur
herpetologists, so it is a delight for us to dignify his ef-
forts towards this aim. The specific name synaptor, a noun
in apposition, stems from the Greek word “συναψις”
meaning “a connection”. In this case, this noun underlines
the connection of these two kinds of herpetologists.
We suggest the following common names: Boehme’s wolf
Snake (English), Böhmes Wolfszahnnatter (German).
Description of the holotype. Habitus. Body elongate,
somewhat laterally compressed; head flattened anterior-
ly, well distinct from the neck; snout depressed and elon-
gate; nostril oval, large, in the middle of the nasal. Eye
moderate, with a vertically elliptic pupil.
SVL 374 mm; TaL 89 mm; TL 463 mm.
291
New Lycodon from Yunnan
Bonn zoological Bulletin 57 (2): 289–296 ©ZFMK
Fig. 2. Ventral view of preserved holotype of Lycodon syn-
aptor sp. n., BMNH 1905.1.30.63 from Dongchuan, 100 km
north of Kunming, Yunnan Province, People’s Republic of Chi-
na. Photograph by Gernot Vogel.
Fig. 4. Dorsal view of Lycodon fasciatus. CIB 9804, from
Ruili City, Yunnan. Note the irregular bands. Photograph by Ger-
not Vogel.
Fig. 3. Lateral view of the right side of the head of preserved
holotype of Lycodon synaptor sp. n., BMNH 1905.1.30.63 from
Dongchuan, 100 km north of Kunming, Yunnan Province,
People’s Republic of China. Photograph by Gernot Vogel.
Dentition. A total of 10 maxillary teeth, with the follow-
ing formula: 4 small anterior teeth + 2 strongly enlarged
teeth + a wide gap + 2 small teeth + a small gap + 2 strong-
ly enlarged, posterior teeth.
Body scalation. 201 VEN (+ 2 preventrals), 68 SC, all
paired. Anal single. Dorsal scales in 17–17–15 rows, the
7 upper rows feebly keeled. Vertebral row not enlarged.
No apical pit detected.
Head scalation. Rostral, triangular, hardly visible from
above; nasal vertically divided by a furrow below and
above the nostril; two small, trapezoidal internasals, wide-
ly in contact each with the other and with prefrontals; two
large prefrontals, longer and wider than internasals; a
rather small ogive-shaped frontal, about 1.3 times longer
than wide and about 0.8 time as long as the suture between
the parietals; 2 large parietals, each edged with three large
scales, 2 upper temporals and a larger paraparietal poste-
riorly; 1 / 1 wide, triangular supraocular; 1 / 1 small pre-
ocular, located above the posterior part of loreal; 2 / 2 pos-
toculars, about the same size; 1 / 1 subrectangular loreal,
elongateand narrow, not entering orbit, in contact with SL
2 and 3, the large preocular, the prefrontal (long contact)
and the posterior part of nasal; 8 / 8 SL, of which the first
four are higher than long, SL 1–2 in contact with nasal,
SL 3–5 entering orbit, 6th and 7th SL largest; 2+2 / 2+2
temporals, lower anterior temporal a bit broader than up-
per one, posterior temporals smaller; 8 / 8 infralabials, IL
1–4 in contact with the first pair of sublinguals; anterior
and posterior pair of sublinguals of about same length, but
anterior pair wider.
Coloration in preservation. Body and tail dark blackish-
brown, with 31 crossbands on body and 9 on tail, narrow
and cream, not speckled; these crossbands, about 1 dor-
sal scale long, widen at their ventrolateral limit, up to about
5–7 dorsal scale long; the first crossband, beginning at the
level of VEN 9; the second crossband is 8 scale rows be-
hind the first one; 9 cream crossbands on the tail, also not
speckled.
The head is uniformly blackish-brown, a broad nuchal col-
lar extends from the 6th and 7th supralabial across the low-
er posterior temporal across the posterior half of the pari-
entals. The underside of the head ist dark in the anterior
half and cream in the posterior one; the throat is cream,
with a dark clowdy spot on the preventrals and the first
ventral.
The venter is dark, with faint cream bands rather regular,
2 ventrals wide and with 3–4 ventrals in between. With-
in these bands some ventrals are dark on one half and
cream on the other, especially in the posterior part of the
body. The under surface of the tail is banded as the ven-
ter with the cream bands about 1.5 SC wide.
292 Gernot Vogel & Patrick David
Bonn zoological Bulletin 57 (2): 289–296 ©ZFMK
Fig. 5. Ventral view of Lycodon fasciatus. CIB 9804, from
Ruili City, Yunnan. Note he whitish colouration of the anterior
part and te speckling of the posterior part. Photograph by Ger-
not Vogel.
Fig. 6. Comparison of lateral head scalation of Lycodon synaptor sp. n. (BM 1905.1.30.63 Holotype) and one L. fasciatus (BNHS
1223) where the Lo touches the eye. This is the case only in 6.3% of all cases seen by us. Please note that in L. synaptor sp. n. the
Lo is well separate from the eye by the preocular scale, whereas it is tapering and narrow in the L. fasciatus, where it is more or
less inserted between 2 sales and falls short from the eye. Drawings by Dick Visser.
Variation. The paratype, MNHN 1905.0283, agrees in
most respects with the description of the holotype with the
throat being dark instead of light. The maxilla are miss-
ing in the paratype. A comparison of the most important
morphological characters is summarized in Table 2.
Distribution. China. Lycodon synaptor sp. n. is current-
ly only known from the region of Dongchuan, Dongchuan
County, in the province of Yunnan, China.
Biology. There is no information available on the biolo-
gy of this species. However, the region of Dongchuan is
highly mountainous. Dongchuan is located between high
mountains of the ranges known as Gongwang Shan and
Liangwang Shan. In the area, the highest point is 4.344
meters high, and lowest is 695 meters asl.
DISCUSSION
Lycodon synaptor sp. n. is superficially similar to L. fas-
ciatus but differs from the whole L. fasciatus group (in-
cluding L. butleri) by the loreal, which does not enter or-
bit in L. synaptor. We have examined six specimens (out
of 35) of Lycodon fasciatus sensu stricto in which the lo-
real does not enter orbit (eight occurrence, three times on
both sides [4.7 %]), but the morphology of the anterior
region of the eye is different. In specimens of L. fascia-
tus in which the loreal does not enter orbit, the posterior
region of this narrow scale is very tapering (Fig. 6). Its
apex is more or less inserted between the preocular and
the 4th SL and falls short from the eye. In contrast, in L.
synaptor, the loreal scale is well separated from the orbit
by a broad preocular. The tail is a bit shorter in L. synap-
tor sp. n. (0.189–0.192 vs. 0.190–0.219 in 29 females of
L. fasciatus). There are also differences in the shape of the
bands and the colouration of the belly (compare Figs 2 and
5). L. synaptor sp. n. has eight infralabials, whereas only
one specimen out of 60 specimens of L. fasciatus had eight
infralabials on both sides and rarely that character is seen
on one side (5 %) in L. fasciatus. L. synaptor sp. n. has
less subcaudals than L. fasciatus (68–69, x = 68.5, versus
70–88, x = 79.9 with one outlier having 67 subcaudals).
For a complete comparison of scale counts see Table 3.
L. synaptor sp. n. differs from the other Chinese and In-
dochinese species as follows: from L. subcinctus by the
fact that in L. synaptor sp. n. both a loreal and a preocu-
lar are present. From L. laoensis, L. zawi and L. capuci-
nus it differs by the anal shield which is single in L. synap-
tor sp. n. but divided in the latter two species. Furthermore
the colouration is much different. From the L. ruhstrati
group it differs by the colouration of the bands (small, reg-
ular, completely light in L. synaptor sp. n., getting broad-
er posteriorly, irregular and partly speckled with brown
in the L. ruhstrati complex), by the number of infralabi-
als (eight in L. synaptor, nine to ten in the L. ruhstrati
group) and by the colour of the belly, which is speckled
or uniform light in the L. ruhstrati group, but clearly band-
ed in L. synaptor sp. n. From L. paucifasciatus Rendahl,
1943, occurring in Vietnam, L. synaptor sp. n. differs by
the number of anterior dorsal scale rows (15–17 in L.
synaptor sp. n. and 19 in L. paucifasciatus).
293
New Lycodon from Yunnan
Bonn zoological Bulletin 57 (2): 289–296 ©ZFMK
Table 2. Pholidosis of the two type type specimens of Lycodon synaptor sp. n.
Characters BMNH 1905.1.30.63 MNHN 1905.0283
holotype paratype
Sex Female Female
SVL 374 395
TaL 87 92
Rel TL 0.192 0.189
ASR 17 15
MSR 17 17
VEN 201 203
SC 68 69
Lo enters eye no no
Dorsal bands 31 30
Tail bands 9 9
First band at VEN no 9 5
Broad base [VEN] 7 3
Broad vertebral [Dorsals] 2 1
Werner (1922) described Dinodon yunnanensis from Yun-
nanfu, now Kunming, Yunnan Province. This species was
synonymised with Lycodon fasciatus by Pope (1935: 188),
but according to our unpublished data, this taxon might
be valid. In any way this name is available, so we com-
pared Lycodon synaptor sp. n. with Dinodon yunnanen-
sis Werner, 1922 for which we re-examined the holotype
(NMW 23417; adult female). Lycodon synaptor sp. n. dif-
fers from D. yunnanensis mainly by the loreal which is
touching the eye in D. yunnanensis, but also by the num-
ber of ventrals (201–203 against 193) and the number of
bands on the body (30–31 vs. 23) and the number of in-
fralabials (eight vs. nine in D. yunnanensis).
While preparing a review for the Lycodon fasciatus group,
we found quite a lot of obviously independent lineages,
294 Gernot Vogel & Patrick David
Bonn zoological Bulletin 57 (2): 289–296 ©ZFMK
Table 2. Important characters in the Lycodon fasciatus / ruhstrati groups.
Lycodon fasciatus synaptor sp. n. ruhstrati futsingensis cardamomensis
n females 35 2 23 6 1
TL, females N=29 N=2 N=22 N=5 N=1
679 487 876 773 545
Rel TL, females 0.190–0.219 0.189–0.192 0.207–0237 0.205–0.217 0.25
N=29 N=2 N=22 N=11 N=1
VEN, females 180–219 201–203 217–229 198–208 223
N=35 N=2 N=23 N=13 N=1
SC, females (67) 70–88 68–69 90–108 78–85 92
N=29 N=2 N=21 N=5 N=1
IL both sexes 9–10 (8)*8 10 (9.11) 9–10 (11) 10
N=120 N=2 N=86 N=44 N=4
Bands 19–43 30–31 19–46 19–33 12–13
N=60 N=2 N=45 N=22 N=4
Tail bands 7–21 9 10–23 9–18 6
N=62 N=2 N=43 N=21 N=4
First band 5–18 5–9 8–17 13–23 unknown
N=51 N=2 N=34 N=18
Broad base 3–12 3 5–9 (12) 5–8.5 unknown
N=51 N=2 N=34 N=18
Lo enters orbit Yes (rarely not**) No No (rarely yes***)No No
N=126 N=4 N=86 N=44 N=4
Belly banded Yes Yes No No No
N=63 N=2 N=25**** N=22 N=4
*8 in 6 cases (5 %), in one specimen from Shillong on both sides (1.6 %)
** not entering in 8 occurrences (6.3 %), three times on both sides (4.7 %) (see above)
*** in 6 specimens, all from Fujian the Lo touched the eye.
**** banded in juveniles only
which seem to constitute distinct species. Some of them
are restricted to small areas. Adiscussion of these lineag-
es will follow in the main review (Vogel & David in prep.).
Lycodon synaptor sp. n. differs so much from other mem-
bers of the group that we decided to describe it separate-
ly. The closest localities we got from Lycodon fasciatus
are from the vicinity of Kunming in Yunnan province,
which lies about 100 km south of Dongchuan. This latter
place (previously Tongchuan or Tongchuan Fu) is the type
locality for several reptile species, as Cuora yunnanensis
(Boulenger 1906), Nanorana yunnanensis (Anderson,
1879), Odorrana grahami (Boulenger 1917) and Bombi-
na maxima (Boulenger 1905). It is a relatively small city
that had about 300.000 inhabitants in 2006.
Revised Key for the genus Lycodon in China
According to our data, there are several unnamed species
of the genus Lycodon living in China. So this should be
regarded as preliminary key.
1. Both a loreal and a preocular scales present ...................... 2
Either loreal or preocular absent ..................... L. subcinctus
2. Anal divided ....................................................................... 3
Anal single .......................................................................... 4
3. Anterior chinshields not more than 1.5 times longer than
posterior ones; loreal in extensive contact with internasal,
when adult no crossbands on body .................. L. capucinus
Anterior chinshields 2 to 3 times longer than posterior ones;
loreal not, or barely in contact with internasal (very rarely a
strong contact), when adult yellow crossbands on forepart
of body ................................................................ L. laoensis
4. Belly banded ....................................................................... 5
Belly not banded ................................................................. 6
5. Lo not touching the eye, 8 lower labials .. L. synaptor sp. n.
Lo usually touching the eye, usually 9–10 lower labials
............................................................................. L. fasciatus
6. Dorsal rows keeled ................................ L. ruhstrati abditus
Dorsal rows smooth ....................................... L. futsingensis
Acknowledgements. The first author is indebted to Annemarie
Ohler and Alain Dubois (MNHN) for the grants to work in the
Paris collection. We are grateful to Silke Schweiger, NMW for
sending pictures of the type of Dinodon yunnanensis. Ke Jiang
and Jian Luo helped a lot with further information on Chinese
Lycodon. Montri Sumontha provided scalecounts. We also thank
Colin J. McCarthy (BMNH), Varad Giri (BNHS), Robert C.
Drewes and Jens V. Vidum (CAS), Wang Yuezhao, Zeng Xiao-
mao and Ermi Zhao (CIB), Alan Resetar (FMNH), Dingqui Rao
(KIZ), Ivan Ineich and Annemarie Ohler (MNHN), Franz Tiede-
mann and Richard Gemel (NMW), Wolfgang Böhme (ZFMK),
Mark-Oliver Rödel and Frank Tillack (ZMB), Frank Glaw and
Dieter Fuchs (ZSM) for the possibility to examine specimens de-
posited in the collection of their respective institutions. Dick Vis-
sers made the drawings for us. Many thanks for that.
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295
New Lycodon from Yunnan
Bonn zoological Bulletin 57 (2): 289–296 ©ZFMK
APPENDIX I
Additional comparative material of Lycodon fasciatus
India. Shillong, Assam, BMNH 92.1.25.1; BMNH
1908.6.23.6; BMNH 1908.6.23.8; BMNH
1907.12.16.28–29; BMNH 94.10.4.2; BNHS 1219–20,
1229.
Laos. Xieng-Khouang, Laos, MNHN 1928.69.
Myanmar. Maymyo, Burma, BMNH 1925.12.22.15–16;
Mogok, Burma, BMNH 1900.9.20.5–7; BMNH
1901.4.26.1Mogkok, Burma, BNHS 1221 “Burma”,
BMNH 1908.6.23.9–11 ; Burma, BNHS 1218; Toungyi,
Shan State, Myanmar, BMNH 91.11.26.31; Bhamo, Bur-
ma, BMNH 1925.4.2.28; Kachin Hills, Burma, BMNH
1925.9.17.10–11; South Shan State, BMNH 1908.6.23.14;
Burma-Siam Border, BMNH 1937.2.1.12; Rangoon, Bur-
ma, BMNH 1940.3.3.3; Maymyo, Burma, BNHS
1222–24; Thandung Hills, BNHS 1228.
People’s Republic of China. Western China CAS 55147;
Yunnan, MNHN 1919.148; “Yunnan Fu” (holotype of
Dinodon yunnanensis), NMW 23417; Kuantun, ZSM
75/1938; Kunming, Yunnan, BMNH 1930.11.16.4; Ruili
City, Yunnan, CIB 9804; RuiLi, Yunnan,, CIB 9805;
XiShuangBanNa, Yunnan, CIB 9806, CIB 9808, CIB
9809; LongChuan GongWa, Yunnan, KIZ 74 I 0035;
LongChuan HuSa, Yunnan, KIZ 74 I 0145; Tengchong
County, Yunnan KIZ 74 II 0262; Menglian, Yunnan, KIZ
75 I 473; TengChong TuanTian, Yunnan, KIZ 74 II 0263;
Kunming, Yunnan, KIZ 73009; Kunming, Yunnan, KIZ
77004; Kunming city, Yunnan, KIZ 83007; Yunnan, KIZ
83017; FMNH 15148; Tibet; MNHN 1921.0465 Tibet;
Yunnan,; ZMB 65453
Thailand. Chiang Mai, Thailand, FMNH 178369; CAS
172715, Southern Thailand ? FMNH 178368, Nan
province Thailand FMNH 270716.
Vietnam. Phong Nha, Vietnam, ZFMK 86448–50 (Gen-
Bank: EU999214-215); ZFMK 80665; Ziegler unreg.
296 Gernot Vogel & Patrick David
Bonn zoological Bulletin 57 (2): 289–296 ©ZFMK
